ISSN 0097-3157
PROCEEDINGS OF THE ACADEMY OF NATURAL SCIENCES OF PHILADELPHIA 158: 1-35
APRIL 2009
A checklist of diatoms from inland waters of the northwestern United States LOREN L. BAHLS Faculty Affiliate, Environmental Studies Program, The University of Montana, 1032 12th Avenue, Helena, MT 59601 Email:
[email protected] This checklist is dedicated to Dr. Ruth Patrick on the occasion of her 101st birthday. ABSTRACT.— A list of diatom taxa from inland waters of the northwestern United States is presented as a product of the author’s bioassessment work over the last 40 years. The list is limited to taxa that have been identified from slides that are deposited in the Montana Diatom Collection. The list includes 1,348 species and infra-species taxa in 118 genera. The list is provided in support of recent efforts to compile a comprehensive and up-to-date diatom flora for the conterminous United States. Slides, acid-cleaned material, geospatial data, and/or ecological data for taxa in the checklist are available from the author.
INTRODUCTION The Northwest region (Figure 1) encompasses all or parts of nine states in the northwestern quarter of the conterminous United States, and includes large portions of the Great Plains, Rocky Mountains, Cascade Mountains, Coast Range, Columbia Plateau, and Great Basin. Diatoms have been subjects of study in the region for over 150 years (Ehrenberg 1849, 1850). The region is noted for its diverse diatom flora and for its rich assortment of endemic species (Sovereign 1958, 1963; Kociolek and Stoermer 1986). Yet, the region remains relatively unexplored for diatoms and underrepresented in major diatom collections (Kociolek 2006). In Montana, diatoms have been used for water quality assessments since 1968 (Bahls et al. 2008). Two products of my bioassessment work in Montana and elsewhere in the Northwest are a collection of diatom slides (the Montana Diatom Collection) and a list of diatom taxa identified from those slides (this checklist). The list is presented here to support recent efforts to compile a comprehensive and up-to-date diatom flora for the conterminous United States (Kociolek 2005, 2006).
MDC will be sent to the Academy of Natural Sciences in Philadelphia. About 3,500 replicate slides, prepared from samples collected since 1999, are housed at the University of Montana Herbarium (MONTU) in Missoula. David Dyer is collections manager for MONTU and the MDC in Missoula.
Fig. 1. Area of the United States represented by samples in the Montana Diatom Collection of Northwest Diatoms and by this checklist.
THE MONTANA DIATOM COLLECTION The Montana Diatom Collection (MDC) of Northwest Diatoms is a collection of permanent strewn mounts of acid-cleaned diatom frustules on glass microscope slides. The MDC contains samples collected from nine states between 1968 and 2008, and now numbers over 12,000 slides representing 8,477 samples from 4,335 discrete sampling sites (Figure 2). At this time the MDC is housed at two locations. The main part of the collection, containing slides made from all samples and acid-cleaned material for about 1,600 samples, is located at my residence in Helena, Montana. Upon my retirement, this part of the
Fig 2. Distribution of samples in the Montana Diatom Collection of Northwest Diatoms by state as of November 11, 2008.
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L.L. BAHLS
About 75% of the sites represented by samples in the MDC are flowing waters, that is, waters labeled “river” or “creek” on USGS topographic maps. The remaining 25% of the sites are other habitats, including lakes and reservoirs, springs, freshwater seeps, saline seeps, mine seeps and pits, wetlands of all sorts, stock ponds, industrial ponds, sewage lagoons, fossil deposits, and soils. About 95% of the samples in the MDC were collected from benthic habitats, almost all of them from natural substrates. Most are composite samples from multiple substrates, but a few are substrate specific. The remaining 5% of the samples are plankton samples, either grab samples or net tows. The MDC is supported by a relational electronic database in Microsoft® Access 2000©. This database allows for linkage of names in the checklist to individual slides and vials of acid-cleaned material in the MDC. Slides and acid-cleaned material are available on loan from the author. The database also contains sample and site attributes, including geospatial data and concurrent environmental data for up to 35 water quality variables. The author welcomes inquires regarding collaboration on research projects involving the biogeography, autecology, taxonomy, and biodiversity of diatoms. THE CHECKLIST The checklist contains 1,348 species and infra-species taxa in 118 genera (Table 1). Names in the checklist are arranged alphabetically by genus and species, followed by author and date of publication. Names, authors and dates have been checked against and corrected to conform with the California Academy of Sciences’ Catalogue of Diatom Names as updated on December 12, 2007. Names in bold face type do not appear in a recent checklist of diatoms from the continental United States (Kociolek 2005). The checklist contains only those taxa that have been confidently identified by qualified analysts; the list does not include 351 unknown taxa that could not be identified with confidence using floras and monographs available to the analysts. Although I made most of the entries to the checklist myself, four other analysts also contributed taxa. Surface waters in the Northwest have an extremely wide range of pH, nutrient, salinity, and temperature values, reflecting the highly varied geology and geography of the region (Bahls 2004). These factors, along with the large longitudinal and latitudinal extent of the region, the region’s relatively small human population, and the region’s many national parks and wilderness areas, greatly contribute to the diversity of diatom species found here. The checklist contains predominantly recent diatoms, but some, e.g., Aulacoseira canadensis, are probably extinct in the region (Bahls et al. 2009).
Although all of the samples on which this checklist is based are from inland waters, this is not a checklist of strictly freshwater diatoms. Many inland waters of the Northwest, especially in the Great Plains and Great Basin, have salinity levels that range from brackish to hypersaline. It is not surprising, therefore, that many of the taxa reported here may be found in a recent monograph on diatoms from marine coasts (Witkowski et al. 2000). Additionally, many species from the Northwest are typical of northern regions and alpine habitats; seventy-one percent of the taxa reported here are also found in Northern Europe (Lange-Bertalot & Steindorf 1996) and 64% are found in the Great Lakes region of North America (Stoermer et al. 1999). Since 2004 I have been preparing autecological and biogeographical profiles for diatom species in the Northwest (Bahls 2004-2006). Each profile includes taxonomic information about the species, references used to identify the species, lists of similar species and frequent associates, one or more photographs of voucher specimens, typical habitat, total number of records in the MDC, distribution maps (state, eco-region, and river basin), and ecological optima (abundance-weighted means) for up to 35 water quality variables. To date, I have prepared profiles for 274 species. Images and referenced voucher slides in existing and future profiles in the Northwest Diatoms series may be consulted to determine my concept of taxa included in this checklist. ACKNOWLEDGMENTS Erich Weber, Bill Cody, Sarah Spaulding, and Dennis Vandermeer contributed taxa to this checklist. Erich Weber, Dennis Vandermeer, Julia Eichman and the Academy of Natural Sciences of Philadelphia have contributed slides to the MDC. Many field workers, too numerous to recall, collected the diatom samples upon which this checklist is based. Dr. Charles Reimer provided training, inspiration, and encouragement for my work. Thanks to my family— Peggy, Charlie, and Brad—for their love and support. Two anonymous reviewers improved the manuscript with their helpful suggestions. I thank Marina Potapova for careful editing of the manuscript, for supporting diatom work in Montana, and for her collaboration and friendship. Although I rarely make mistakes, I take full responsibility for all errors of commission and omission in this publication. This project was funded with a grant from my savings account. LITERATURE CITED Bahls, L. L. 2004-2006. Northwest diatoms: a photographic
A CHECKLIST OF DIATOMS FROM INLAND WATERS OF THE NORTHWESTERN U.S. catalogue of species in the Montana Diatom Collection, with ecological optima, associates, and distribution records for the nine northwestern United States, Volumes 1-3. Hannaea, Helena, Montana. Bahls, L., M. Potapova, M.-A. Fallu, and R. Pienitz. 2009. Aulacoseira canadensis and Aulacoseira crassipunctata (Bacillariophyta) in North America. Nova Hedwigia Beiheft 135:167-184. Bahls, L. L., M. Teply, R. Sada de Suplee, and M. W. Suplee. 2008. Note: Diatom biocriteria development and water quality assessment in Montana: A brief history and status report. Diatom Research 23(2):533-540. Ehrenberg, C. G. 1849. Über das mächtigste bis jetzt bekannt gewordene (angeblich 500 Fufs mächtige) Lager von mikroskopischen reinen kieselalgen Süsswasser-Formen am Wasserfall-Flusse in Oregon. Bericht über die zur Bekanntmachung geeingneten Verhandlungen der Königlichen Pruessischen Akademie der Wissenschaften zu Berlin 1849:76-87. Ehrenberg, G. C. 1850. On infusorial deposits on the River Chutes in Oregon. American Journal of Science (Second Series) 9:140. Kociolek, J. P. 2005. A checklist and preliminary bibliography of the recent, freshwater diatoms of inland environments of the continental United States. Proceedings of the California Academy of Sciences (Fourth Series) 56(27):395-525. Kociolek, J. P. 2006. Some thoughts on the development
of a diatom flora for freshwater ecosystems in the continental United States and a listing of recent taxa described from U. S. freshwaters. Proceedings of the California Academy of Sciences (Fourth Series) 57(21):561-586. Kociolek, J. P., and E. F. Stoermer. 1986. Observations on North American Gomphoneis (Bacillariophyceae). II. Descriptions and ultrastructure of two new species. Transactions of the American Microscopical Society 105(2):141-151. Lange-Bertalot, H., and A. Steindorf. 1996. Rote liste der limnischen kieselalgen (Bacillariophyceae) Deutschlands. Schr.-R. f. Vegetationskde. 28:633-677. Sovereign, H. E. 1958. The diatoms of Crater Lake, Oregon. Transactions of the American Microscopical Society 77(2):96-134. Sovereign, H. E. 1963. New and rare diatoms from Oregon and Washington. Proceedings of the California Academy of Sciences (Fourth Series) 31(14):349-368. Stoermer, E. F., R. G. Kreis, and N. A. Andresen. 1999. Checklist of diatoms from the Laurentian Great Lakes. II. Journal of Great Lakes Research 25(3):515-566. Witkowski, A., H. Lange-Bertalot, and D. Metzeltin. 2000. Diatom Flora of Marine Coasts I. Volume 7 in LangeBertalot, H. (ed.), Diversity-Taxonomy-Identification, Iconographia Diatomologica: Annotated Diatom Micrographs. A. R. G. Gantner Verlag K. G., Ruggell, Liechtenstein.
Table 1. Distribution of species and infra-species taxa among genera in this checklist.
Genus Achnanthes Achnanthidium Actinocyclus Actinoptychus Adlafia Amphipleura Amphora Aneumastus Anomoeoneis Asterionella Aulacoseira Bacillaria Berkeleya Biremis Brachysira Caloneis Campylodiscus Capartogramma Cavinula Chaetoceros
Author and Date of Publication Bory, 1822 Kützing, 1844 Ehrenberg, 1837 Ehrenberg, 1843 Lange-Bertalot, 1998 Kützing, 1844 Ehrenberg, 1844 Mann & Stickle, 1990 E. Pfitzer, 1871 Hassall, 1850 Thwaites, 1848 Gmelin, 1791 Greville, 1827 Mann & Cox, 1990 Kützing, 1836 Cleve, 1894 Ehrenberg, 1844 Kufferath, 1956 Mann & Stickle, 1990 Ehrenberg, 1844
3
Number of Taxa 28 19 1 1 4 2 24 2 3 1 17 1 1 3 8 18 1 1 6 2
4 Table 1. cont.
Chamaepinnularia Cocconeis Craticula Ctenophora Cyclostephanos Cyclotella Cylindrotheca Cymatopleura Cymbella Cymbellonitzschia Cymbopleura Decussata Denticula Diadesmis Diatoma Diatomella Dickieia Didymosphenia Diploneis Discostella Distrionella Ellerbeckia Encyonema Encyonopsis Entomoneis Eolimna Epithemia Eucocconeis Eunotia Fallacia Fistulifera Fragilaria Fragilariforma Frustulia Geissleria Germainiella Gomphoneis Gomphonema Gomphosphenia Gyrosigma Hannaea Hantzschia Haslea Hippodonta Hydrosera Karayevia Kobayasiella Lemnicola Luticola Mastogloia
L.L. BAHLS
Lange-Bertalot & Krammer, 1996 Ehrenberg, 1837 Grunow, 1867 (Grunow) Williams & Round, 1986 Round, 1987 (Kützing) Brébisson, 1838 Rabenhorst, 1859 Smith, 1851 Agardh, 1830 Hustedt, 1924 Krammer, 1999 (Ehrenberg) Lange-Bertalot & Metzeltin, 2000 Kützing, 1844 Kützing, 1844 Bory, 1824 Greville, 1855 Berkeley, 1844 Schmidt, 1899 Ehrenberg, 1844 Houk & Klee, 2004 Williams, 1990 Crawford, 1988 Kützing, 1833 Krammer, 1997 Ehrenberg, 1845 Lange-Bertalot & Schiller, 1997 Brébisson, 1838 Cleve, 1912 Ehrenberg, 1837 Stickle & Mann, 1990 Lange-Bertalot, 1997 Lyngbye, 1819 Williams & Round, 1988 Rabenhorst, 1853 Lange-Bertalot & Metzeltin, 1996 (Germain) Lange-Bertalot & Metzeltin, 2005 Cleve, 1894 Ehrenberg, 1832 Lange-Bertalot, 1995 Hassall, 1845 Patrick, 1966 Grunow, 1877 Simonsen, 1974 Lange-Bertalot et al., 1996 Wallich, 1858 Round & Bukhtiyarova, 1998 (Cleve) Lange-Bertalot, 1998 Round & Basson, 1997 Mann, 1990 Thwaites, 1856
6 11 20 1 3 14 2 3 54 1 26 1 6 6 6 1 1 1 16 4 1 1 49 18 4 1 8 5 38 8 1 16 4 11 9 1 8 99 1 11 2 10 1 6 1 5 1 1 9 6
A CHECKLIST OF DIATOMS FROM INLAND WATERS OF THE NORTHWESTERN U.S. Table 1. cont.
Mayamaea Melosira Meridion Microcostatus Muelleria Navicula Neidiopsis Neidium Nitzschia Nupela Opephora Orthoseira Parlibellus Pinnularia Placoneis Plagiotropis Planothidium Platessa Pleurosigma Pleurosira Proshkinia Psammothidium Pseudostaurosira Puncticulata Reimeria Rhoicosphenia Rhopalodia Rossithidium Scoliopleura Sellaphora Simonsenia Skeletonema Stauroforma Stauroneis Staurosira Staurosirella Stenopterobia Stephanocostis Stephanodiscus Surirella Synedra Tabellaria Tabularia Terpsinoe Tetracyclus Thalassiosira Tryblionella Urosolenia Total Taxa
Lange-Bertalot, 1997 Agardh, 1824 Agardh, 1824 Johansen & Sray, 1998 (Frenguelli) Frenguelli, 1945 Bory, 1822 Lange-Bertalot & Metzeltin, 1999 Pfitzer, 1871 Hassall, 1845 Vyvermann & Compére, 1991 Petit, 1888 Thwaites, 1848 Cox, 1988 Ehrenberg, 1843 Mereschkowsky, 1903 Pfitzer, 1871 Round & Bukhtiyarova, 1996 Lange-Bertalot, 2004 Smith, 1852 (Meneghini) Trevisan di San Leon, 1848 Karayeva, 1978 Bukhtiyarova & Round, 1996 Williams & Round, 1988 Håkansson, 2002 Kociolek & Stoermer, 1987 Grunow, 1860 Müller, 1895 Round & Bukhtiyarova, 1996 Grunow, 1860 Mereschkowsky, 1902 Lange-Bertalot, 1979 Greville, 1865 Flower et al., 1996 Ehrenberg, 1843 Ehrenberg, 1843 Williams & Round, 1988 Brébisson, 1896 Genkal & Kuzmina, 1985 Ehrenberg, 1845 Turpin, 1828 Ehrenberg, 1830 Ehrenberg, 1844 (Kützing) Williams & Round, 1986 Ehrenberg, 1843 Ralfs, 1843 Cleve, 1873 W. Smith, 1853 Round & Crawford, 1990
5
6 1 3 3 1 209 2 19 133 4 3 2 2 68 9 3 22 3 4 1 2 17 7 5 3 1 8 3 1 14 1 2 1 29 4 11 3 1 8 23 20 3 2 1 1 4 12 1 1,348
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L.L. BAHLS
Checklist Achnanthes acares Hohn & Hellerman, 1963 Achnanthes bahusiensis (Grunow) Lange-Bertalot in Lange-Bertalot & Krammer, 1989 Achnanthes brevipes Agardh, 1824 Achnanthes coarctata (Brébisson ex W. Smith) Grunow in Cleve & Grunow, 1880 Achnanthes curtissima Carter, 1963 Achnanthes hintzii Lange-Bertalot & Krammer, 1989 Achnanthes holstii Cleve, 1881 Achnanthes imperfecta Schimanski, 1978 Achnanthes impexiformis Lange-Bertalot in Lange-Bertalot & Krammer, 1989 Achnanthes kriegeri Krasske, 1943 Achnanthes lemmermanni Hustedt, 1933 Achnanthes levanderi Hustedt, 1933 Achnanthes linearis f. curta H. L. Smith ex Boyer, 1916 Achnanthes minuscula Hustedt, 1945 Achnanthes nitidiformis Lange-Bertalot in Lange-Bertalot & Krammer, 1991 Achnanthes nodosa Cleve, 1900 Achnanthes oblongella Østrup, 1902 Achnanthes pericava Carter, 1966 Achnanthes pseudoswazi Carter, 1963 Achnanthes ricula Hohn & Hellerman, 1963 Achnanthes rostellata Cleve-Euler, 1934 Achnanthes subexigua Hustedt, 1934 Achnanthes subhudsonis Hustedt, 1921 Achnanthes subhudsonis var. kraeuselii (Cholnoky) Cholnoky, 1954 Achnanthes subsalsa Petersen, 1928 Achnanthes taeniata Grunow in Cleve & Grunow, 1880 Achnanthes thermalis (Rabenhorst) Schönfeldt, 1907 Achnanthes ziegleri Lange-Bertalot, 1993
MT, OR MT ID CA, MT ID, OR, WY ID, MT, OR MT MT ID, MT, WA CA, MT, OR, WA CA, WY MT MT MT ID, MT, OR MT, OR, WA, WY CA, ID, MT, OR, WA CA OR OR MT NV CA, ID, MT, OR, WA CA ID, MT CA, MT MT, SD, WA MT
Achnanthidium affine (Grunow) Czarnecki, 1994 Achnanthidium catenatum (Bily & Marvan) Lange-Bertalot, 1999 Achnanthidium deflexum (Reimer) Kingston, 2000 Achnanthidium duthii (Sreenivasa) Edlund, 1994 Achnanthidium eutrophilum (Lange-Bertalot in Lange-Bertalot & Metzeltin) Lange-Bertalot, 1999 Achnanthidium exiguum (Grunow) Czarnecki, 1994 Achnanthidium exiguum var. constrictum (Grunow) Andresen, Stoermer & Kreis, 2000 Achnanthidium exile (Kützing) Round & Bukhtiyarova, 1996 Achnanthidium gracillimum (Meister) Lange-Bertalot, 2004 Achnanthidium kranzii (Lange-Bertalot) Round & Bukhtiyarova, 1996 Achnanthidium microcephalum Kützing, 1844 Achnanthidium minutissimum (Kützing) Czarnecki, 1994 Achnanthidium minutissimum var. jackii (Rabenhorst) Lange-Bertalot, 1980 Achnanthidium neomicrocephalum Lange-Bertalot & Staab, 2004 Achnanthidium pyrenaicum (Hustedt) Kobayasi, 1997 Achnanthidium rivulare Potapova & Ponader, 2004 Achnanthidium saprophilum (Kobayasi & Mayama) Round & Bukhtiyarova, 1996 Achnanthidium semiapertum (Guermeur) Andresen, Stoermer & Kreis, 2000 Achnanthidium thienemannii (Hustedt) Lange-Bertalot, 2004
CA, ID, MT, OR, SD, WA, WY CA, ID, MT, WA CA, ID, MT, OR, SD, WA, WY CA, MT CA CA, ID, MT, ND, OR, SD, WA, WY MT WA MT MT ID, MT CA, ID, MT, ND, NV, OR, SD, WA, WY MT ID, MT CA, ID, MT, OR, WA, WY CA, ID, MT, OR, WA, WY MT, OR MT MT, WY
Actinocyclus normanii (Gregory ex Greville) Hustedt, 1957
CA
A CHECKLIST OF DIATOMS FROM INLAND WATERS OF THE NORTHWESTERN U.S.
7
Checklist cont. Actinoptychus senarius (Ehrenberg) Ehrenberg, 1843
CA
Adlafia bryophila (Petersen) Moser, Lange-Bertalot & Metzeltin, 1998 Adlafia minuscula (Grunow) Lange-Bertalot in Lange-Bertalot & Genkal, 1999 Adlafia minuscula var. muralis (Grunow) Lange-Bertalot in Lange-Bertalot & Genkal, 1999 Adlafia suchlandtii (Hustedt) Moser, Lange-Bertalot & Metzeltin, 1998
CA, ID, MT, ND, OR, SD, WA, WY CA, ID, MT, ND, OR, SD, WA, WY CA, ID, MT, ND, SD, WY CA, ID, MT, OR, WA, WY
Amphipleura kriegeriana (Krasske) Hustedt, 1954 Amphipleura pellucida (Kützing) Kützing, 1844
MT CA, ID, MT, ND, NV, OR, SD, WA, WY
Amphora aequalis Krammer, 1980 Amphora coffeaeformis (Agardh) Kützing, 1844 Amphora coffeaformis var. acutiuscula (Kützing) Rabenhorst, 1864 Amphora copulata (Kützing) Schoeman & Archibald, 1986 Amphora delicatissima Krasske in Hustedt, 1930 Amphora dusenii Brun, 1901 Amphora fogediana Krammer in Krammer & Lange-Bertalot, 1985 Amphora hassiaca Krammer & Strecker in Krammer, 1997 Amphora holsatica Hustedt, 1930 Amphora holsaticoides Nagumo & Kobayasi, 1990 Amphora inariensis Krammer, 1980 Amphora montana Krasske, 1932 Amphora normannii Rabenhorst, 1864 Amphora ovalis (Kützing) Kützing, 1844 Amphora pediculus (Kützing) Grunow in Schmidt et al., 1875 Amphora pseudoholsatica Nagumo & Kobayasi, 1990 Amphora schroederi Hustedt, 1921 Amphora spitzbergensis VanLandingham, 1967 Amphora strigosa Hustedt, 1949 Amphora tenerrima Aleem & Hustedt, 1951 Amphora tenuissima Hustedt, 1955 Amphora thumensis (Mayer) Cleve-Euler, 1932 Amphora tumida Hustedt, 1956 Amphora veneta Kützing, 1844
MT CA, ID, MT, SD, WY MT CA, ID, MT, ND, OR, SD, WY MT, ND, SD, WY CA, ID, MT, SD, WY ID, MT, OR, WA WY MT, SD, WY MT CA, ID, MT, NV, OR, SD, WA, WY CA, ID, MT, ND, OR, SD, WA, WY ND, SD CA, ID, MT, OR, WY CA, ID, MT, ND, NV, OR, SD, WA, WY MT WA MT WY MT MT MT SD CA, ID, MT, ND, NV, OR, SD, WA, WY
Aneumastus minor Lange-Bertalot, 1993 Aneumastus tusculus (Ehrenberg) Mann & Stickle in Round, Crawford & Mann, 1990
MT, WY ID, MT, OR, WY
Anomoeoneis costata (Kützing) Hustedt, 1959 Anomoeoneis sculpta (Ehrenberg) Cleve, 1895 Anomoeoneis sphaerophora Pfitzer, 1871
CA, MT CA, MT CA, ID, MT, ND, NV, OR, WY
Asterionella formosa Hassall, 1850
CA, ID, MT, OR, WA, WY
Aulacoseira alpigena (Grunow) Krammer, 1991 Aulacoseira ambigua (Grunow) Simonsen, 1979 Aulacoseira canadensis (Hustedt) Simonsen, 1979 Aulacoseira crenulata (Ehrenberg) Thwaites, 1848 Aulacoseira distans (Ehrenberg) Simonsen, 1979 Aulacoseira granulata (Ehrenberg) Simonsen, 1979
CA, ID, MT, OR, SD, WA, WY CA, ID, MT, ND, OR CA, ID, MT, ND, OR, SD, WA, WY CA, ID, MT, OR, WY CA, ID, MT, ND, OR, WA, WY CA, ID, MT, ND, OR, SD, WA, WY
8
L.L. BAHLS
Checklist cont. Aulacoseira granulata var. angustissima (O. Müller) Simonsen, 1979 Aulacoseira islandica (O. Müller) Simonsen, 1979 Aulacoseira italica (Ehrenberg) Simonsen, 1979 Aulacoseira lacustris (Grunow in Van Heurck) Krammer, 1991 Aulacoseira laevissima (Grunow in Van Heurck) Krammer, 1991 Aulacoseira lirata (Ehrenberg) Ross in Hartley, 1986 Aulacoseira pfaffiana (Reinsch) Krammer, 1991 Aulacoseira subarctica (O. Müller) Haworth, 1990 Aulacoseira tenuior (Grunow) Krammer, 1991 Aulacoseira tethra Haworth, 1988 Aulacoseira valida (Grunow in Van Heurck) Krammer, 1991
CA, ID, MT, ND, OR, SD, WA, WY CA, MT, ND, OR, WA, WY CA, ID, MT, ND, NV, OR, SD, WA, WY CA, MT MT CA, MT, OR ID, MT MT CA, MT, OR MT MT, OR, WY
Bacillaria paradoxa Gmelin in Linneaeus, 1788
CA, MT, ND, SD, WY
Berkeleya rutilans (Trentepohl ex Roth) Grunow, 1880
WY
Biremis circumtexta (Meister ex Hustedt) Lange-Bertalot & Witkowski in Witkowski, LangeBertalot & Metzeltin, 2000 Biremis motschii (Meister) Lange-Bertalot in Werum & Lange-Bertalot, 2004 Biremis zachariasii (Reichelt) Edlund, Andresen & Soninkhishig, 2001
MT, ND, SD, WY MT ID, MT
Brachysira brebissonii Ross in Hartley, 1986 Brachysira microcephala (Kützing) Compére, 1986 Brachysira neoexilis Lange-Bertalot in Lange-Bertalot & Moser, 1994 Brachysira procera Lange-Bertalot & Moser, 1994 Brachysira serians (Brébisson) Round & Mann, 1981 Brachysira styriaca (Grunow in Van Heurck) Ross in Hartley, 1986 Brachysira vitrea (Grunow) Ross in Hartley, 1986 Brachysira zellensis (Grunow) Round & Mann, 1981
CA, MT, OR CA, ID, MT, ND, SD, WA, WY CA, MT MT MT MT CA, ID, MT, WA MT
Caloneis alpestris (Grunow) Cleve, 1894 Caloneis amphisbaena (Bory) Cleve, 1894 Caloneis bacillum (Grunow) Cleve, 1894 Caloneis budensis (Grunow) Krammer in Krammer & Lange-Bertalot, 1985 Caloneis hyalina Hustedt, 1937 Caloneis lauta Carter & Bailey-Watts, 1981 Caloneis leptosoma (Grunow) Krammer in Krammer & Lange-Bertalot, 1985 Caloneis molaris (Grunow) Krammer in Krammer & Lange-Bertalot, 1985 Caloneis obtusa (W. Smith) Cleve, 1894 Caloneis oregonica (Ehrenberg) Patrick in Patrick & Reimer, 1966 Caloneis permagna (Bailey) Cleve, 1894 Caloneis pulchra Messikommer, 1927 Caloneis schumanniana (Grunow in Van Heurck) Cleve, 1894 Caloneis silicula (Ehrenberg) Cleve, 1894 Caloneis tenuis (Gregory) Krammer in Krammer & Lange-Bertalot, 1985 Caloneis thermalis (Grunow) Krammer in Krammer & Lange-Bertalot, 1985 Caloneis undulata (Gregory) Krammer in Krammer & Lange-Bertalot, 1985 Caloneis westii (W. Smith) Hendey, 1964
ID, MT CA, ID, MT, ND, OR, SD, WY CA, ID, MT, ND, NV, OR, SD, WA, WY MT MT MT, WY ID, MT CA, ID, MT, ND, OR, SD, WY MT MT CA, SD CA, MT, OR CA, ID, MT, ND, OR, SD, WY CA, ID, MT, ND, NV, OR, SD, WA, WY CA, ID, MT, NV, OR, SD, WA, WY MT ID, MT, OR, WA CA, MT
Campylodiscus hibernicus Ehrenberg, 1845
CA, ID, MT, SD
A CHECKLIST OF DIATOMS FROM INLAND WATERS OF THE NORTHWESTERN U.S.
9
Checklist cont. Capartogramma crucicula (Grunow ex Cleve) Ross, 1963
CA
Cavinula cocconeiformis (Gregory ex Greville) Mann & Stickle in Round, Crawford & Mann, 1990 Cavinula jaernefeltii (Hustedt) Mann & Stickle in Round, Crawford & Mann, 1990 Cavinula lapidosa (Krasske) Lange-Bertalot in Lange-Bertalot & Metzeltin, 1996 Cavinula pseudoscutiformis (Hustedt) Mann & Stickle in Round, Crawford & Mann, 1990 Cavinula scutelloides (W. Smith) Lange-Bertalot in Lange-Bertalot & Metzeltin, 1996 Cavinula scutiformis (Grunow ex Schmidt) Mann & Stickle in Round, Crawford & Mann, 1990
CA, ID, MT, OR, WA, WY ID, MT, OR ID, MT, OR ID, MT, OR, WY ID, MT MT
Chaetoceros elmorei Boyer, 1914 Chaetoceros muelleri Lemmermann, 1898
MT CA, MT, ND, WY
Chamaepinnularia bremensis (Hustedt) Lange-Bertalot in Lange-Bertalot & Metzeltin, 1996 Chamaepinnularia evanida (Hustedt) Lange-Bertalot in Lange-Bertalot & Metzeltin, 1996 Chamaepinnularia mediocris (Krasske) Lange-Bertalot in Lange-Bertalot & Metzeltin, 1996 Chamaepinnularia soehrensis (Krasske) Lange-Bertalot & Krammer in Lange-Bertalot & Metzeltin, 1996 Chamaepinnularia soehrensis var. hassiaca (Krasske) Lange-Bertalot in Lange-Bertalot & Metzeltin, 1996 Chamaepinnularia soehrensis var. muscicola (Petersen) Lange-Bertalot & Krammer in LangeBertalot & Metzeltin, 1996
CA, ID, MT, NV, OR, WA, WY CA, ID, MT, WA MT
Cocconeis disculus (Schumann) Cleve, 1882 Cocconeis klamathensis Sovereign, 1958 Cocconeis neodiminuta Krammer, 1990 Cocconeis neothumensis Krammer, 1990 Cocconeis pediculus Ehrenberg, 1838 Cocconeis placentula Ehrenberg, 1838 Cocconeis placentula var. euglypta (Ehrenberg) Grunow, 1884 Cocconeis placentula var. lineata (Ehrenberg) Van Heurck, 1885 Cocconeis pseudothumensis Reichardt, 1982 Cocconeis rugosa Sovereign, 1958 Cocconeis scutellum Ehrenberg, 1838
ID, MT, OR, WA OR CA, ID, MT, OR, SD, WA, WY CA, ID, MT, OR, WA, WY CA, ID, MT, OR, ND, NV, SD, WA, WY CA, ID, MT, OR, ND, NV, SD, WA, WY CA, ID, MT, OR, ND, NV, SD, WA, WY CA, ID, MT, OR, ND, NV, SD, WA, WY CA, MT, OR, WA CA, OR OR
Craticula accomoda (Hustedt) Mann in Round, Crawford & Mann, 1990 Craticula accomodiformis Lange-Bertalot, 1993 Craticula acidoclinata Lange-Bertalot & Metzeltin, 1996 Craticula ambigua (Ehrenberg) Mann in Round, Crawford & Mann, 1990 Craticula buderi (Hustedt) Lange-Bertalot in Rumrich, Lange-Bertalot & Rumrich, 2000 Craticula citrus (Krasske) Reichardt, 1997 Craticula cuspidata (Kützing) Mann in Round, Crawford & Mann, 1990 Craticula dissociata (Reichardt) Reichardt, 1997 Craticula elkab (O. Müller) Lange-Bertalot, Kubser & Cocquyt in Kubser & Cocquyt, 2007 Craticula halopannonica Lange-Bertalot, 2001 Craticula halophila (Grunow ex Van Heurck) Mann in Round, Crawford & Mann, 1990 Craticula halophilioides (Hustedt) Lange-Bertalot, 2001 Craticula minusculoides (Hustedt) Lange-Bertalot, 2001 Craticula molesta (Krasske) Lange-Bertalot & Willmann in Lange-Bertalot, Kulbs, Lauser, NorpelSchempp & Willmann, 1996 Craticula molestiformis (Hustedt) Lange-Bertalot in Rumrich, Lange-Bertalot & Rumrich, 2000 Craticula riparia (Hustedt) Lange-Bertalot, 1993
CA, ID, MT, NV, OR, SD, WY MT ID, MT CA, ID, MT, ND, OR, SD, WY CA, MT, ND, OR, SD, WY CA, MT CA, ID, MT, ND, NV, OR, SD, WY ID MT CA, MT ID, MT, SD, WY MT, SD ID, MT
CA, ID, MT MT ID, MT
MT CA, MT, WY CA, MT
10
L.L. BAHLS
Checklist cont. Craticula silviae Lange-Bertalot 2001 Craticula submolesta (Hustedt) Lange-Bertalot in Lange-Bertalot & Metzeltin, 1996 Craticula vixnegligenda Lange-Bertalot, 1993 Craticula vixvisibilis (Hustedt) Lange-Bertalot, 1993
CA, MT MT MT MT
Ctenophora pulchella (Ralfs ex Kützing) Williams & Round, 1986
CA, MT, ND, SD, WY
Cyclostephanos dubius (Fricke) Round validated by Theriot, Stoermer & Håkansson, 1987 Cyclostephanos invisitatus (Hohn & Hellerman) Theriot, Stoermer & Håkansson, 1987 Cyclostephanos tholiformis Stoermer, Håkansson & Theriot, 1987
ID, MT CA, ID, MT, ND, OR, SD, WA, WY CA, MT, SD, WY
Cyclotella antiqua W. Smith, 1853 Cyclotella atomus Hustedt, 1937 Cyclotella comensis Grunow in Van Heurck, 1882 Cyclotella cyclopuncta Håkansson & Carter, 1990 Cyclotella delicatula Hustedt, 1952 Cyclotella distinguenda Hustedt, 1928 Cyclotella gamma Sovereign, 1963 Cyclotella hakanssoniae Wendker, 1991 Cyclotella kuetzingiana Thwaites, 1848 Cyclotella meneghiniana Kützing, 1844 Cyclotella michiganiana Skvortzow, 1937 Cyclotella ocellata Pantocsek, 1902 Cyclotella striata (Kützing) Grunow in Cleve & Grunow, 1880 Cyclotella tripartita Håkansson, 1990
MT ID, MT, ND, OR, SD, WA, WY ID, MT, OR MT MT ID, MT MT MT MT CA, ID, MT, ND, NV, OR, SD, WA, WY ID, MT CA, ID, MT, OR, WY MT CA, MT
Cylindrotheca closterium (Ehrenberg) Reiman & Lewin, 1964 Cylindrotheca gracilis (Brébisson ex Kützing) Grunow in Van Heurck, 1882
MT, ND, SD, WY ID, MT, ND, SD, WY
Cymatopleura elliptica (Brébisson ex Kützing) W. Smith, 1851 Cymatopleura solea (Brébisson in Brébisson & Godey) W. Smith, 1851 Cymatopleura solea var. apiculata (W. Smith) Ralfs in Pritchard, 1861
CA, ID, MT, ND, NV, OR, SD, WY CA, ID, MT, ND, NV, OR, SD, WA, WY ID, MT
Cymbella affiniformis Krammer, 2002 Cymbella affinis Kützing, 1844 Cymbella alpestris Krammer, 2002 Cymbella aspera (Ehrenberg), 1894 Cymbella cleve-eulerae Krammer, 2002 Cymbella cymbiformis Agardh, 1830 Cymbella delicatula Kützing, 1849 Cymbella diluviana (Krasske) Florin, 1971 Cymbella diversiformis Krammer, 2002 Cymbella diversistigmata Krammer, 2002 Cymbella dorsenotata Østrup, 1910 Cymbella excisa Kützing, 1844 Cymbella excisiformis Krammer, 2002 Cymbella frigida Cleve-Euler, 1939 Cymbella graciliformis Krammer, 2002 Cymbella hantzschiana (Rabenhorst) Krammer, 2002 Cymbella hustedtii Krasske, 1923
ID, MT, WY CA, ID, MT, ND, NV, SD, WA, WY MT CA, ID, MT WY CA, ID, MT, ND, OR, SD, WA, WY CA, ID, MT, SD, WA, WY ID, MT MT MT, WA ND CA, ID, MT, ND, NV, OR, SD, WA, WY ID, MT, WY MT MT CA, MT, ND, SD ID, MT, ND, SD, WA
A CHECKLIST OF DIATOMS FROM INLAND WATERS OF THE NORTHWESTERN U.S.
11
Checklist cont. Cymbella hybrida Grunow in Cleve & Möller, 1878 Cymbella janischii ( Schmidt in Schmidt et al.) Cleve, 1894 Cymbella kappii (Cholnoky) Cholnoky, 1956 Cymbella laevis Naegeli in Kützing, 1849 Cymbella laevis var. lata Krammer, 2002 Cymbella lanceolata (Agardh) Agardh, 1830 Cymbella maggiana Krammer, 2002 Cymbella mexicana (Ehrenberg) Cleve, 1894 Cymbella neocistula Krammer, 2002 Cymbella neocistula var. lunata Krammer, 2002 Cymbella neogena (Grunow) Krammer, 2002 Cymbella neoleptoceros Krammer, 2002 Cymbella obtusiformis Krammer, 2002 Cymbella parviformis Krammer, 2002 Cymbella percymbiformis Krammer, 2002 Cymbella perparva Krammer, 2002 Cymbella pervarians Krammer, 2002 Cymbella proxima Reimer in Patrick & Reimer, 1975 Cymbella pusilla Grunow in Schmidt et al., 1875 Cymbella rainierensis Sovereign, 1963 Cymbella reinhardtii Grunow in Schmidt et al., 1875 Cymbella rumrichae Krammer, 2002 Cymbella schimanskii Krammer, 1982 Cymbella schimanskii var. excelsa (Meister) Krammer, 2002 Cymbella simonsenii Krammer in Krammer & Lange-Bertalot, 1985 Cymbella stigmaphora Østrup, 1910 Cymbella strontiana Krammer, 2002 Cymbella subarctica Krammer, 2002 Cymbella subcistula Krammer, 2002 Cymbella subhelvetica Krammer, 2002 Cymbella subleptoceros Krammer, 2002 Cymbella subturgidula Krammer, 2002 Cymbella transilvanica Krammer, 2002 Cymbella tristis Krasske, 1939 Cymbella tumida (Brébisson in Kützing) Van Heurck, 1882-1885 Cymbella turgidula Grunow in Schmidt et al., 1875 Cymbella vulgata Krammer, 2002
MT CA, ID, MT, OR, WA, WY MT MT, OR, SD, WY MT CA, MT, ND, WA CA, MT, OR CA, ID, MT, ND, OR, SD, WA, WY CA, ID, MT, ND, OR, SD, WA, WY MT MT ID, MT, ND, SD MT MT, WY CA, MT CA, ID, MT, WY MT, ND, SD CA, ID, MT, ND, WY CA, MT, ND, SD, WY MT MT MT MT MT CA MT MT MT MT MT MT, SD, WY MT, ID, MT, ND, NV, OR, WA, WY ND MT CA, ID, MT, NV, OR, WA, WY ID, MT, OR, SD CA, MT, SD, WY
Cymbellonitzschia diluviana Hustedt, 1950
CA, ID, MT, OR, WY
Cymbopleura amphicephala (Naegeli) Krammer, 2003 Cymbopleura anglica (Lagerstedt) Krammer, 2003 Cymbopleura angustata (W. Smith) Krammer, 2003 Cymbopleura apiculata Krammer, 2003 Cymbopleura austriaca (Grunow) Krammer, 2003 Cymbopleura crassipunctata Krammer, 2003 Cymbopleura cuspidata (Kützing) Krammer, 2003 Cymbopleura frequens Krammer, 2003 Cymbopleura hauckii (Van Heurck) Krammer, 2003 Cymbopleura heinii Lange-Bertalot & Krammer, 1999
CA, ID, MT, ND, SD, WY MT MT ID, MT MT, SD ID ID, MT MT MT CA, MT
12
L.L. BAHLS
Checklist cont. Cymbopleura hercynica (Schmidt) Krammer, 2003 Cymbopleura hybrida var. capitata (Grunow) Krammer, 2003 Cymbopleura inaequalis (Ehrenberg) Krammer, 2003 Cymbopleura incerta (Grunow) Krammer, 2003 Cymbopleura incertiformis Krammer, 2003 Cymbopleura lapponica (Grunow) Krammer, 2003 Cymbopleura lata (Grunow) Krammer, 2003 Cymbopleura naviculiformis (Auerswald ex Heiberg) Krammer, 2003 Cymbopleura oblongata Krammer, 2003 Cymbopleura rhomboidea Krammer, 2003 Cymbopleura rupicola (Grunow) Krammer, 2003 Cymbopleura subaequalis (Grunow) Krammer, 2003 Cymbopleura subaequalis var. truncata Krammer, 2000 Cymbopleura subcuspidata (Krammer) Krammer, 2003 Cymbopleura sublanceolata Krammer, 2003 Cymbopleura subrostrata (Cleve) Krammer, 2003
MT MT MT MT MT MT MT CA, ID, MT, ND, NV, OR, SD, WY, WY MT WY MT, ND ID, MT, WY MT ID, MT MT MT, OR, WY
Decussata placenta (Ehrenberg) Lange-Bertalot, 2000
MT, OR, WY
Denticula elegans Kützing, 1844 Denticula eximia Krammer & Lange-Bertalot in Lange-Bertalot & Krammer, 1987 Denticula kuetzingii Grunow, 1862 Denticula subtilis Grunow, 1862 Denticula tenuis Kützing, 1844 Denticula valida (Pedicino) Grunow in Van Heurck, 1882-1885
ID, MT, OR, SD, WY MT CA, ID, MT, ND, OR, SD, WY CA, ID, MT, ND, OR, SD, WY CA, ID, MT, WY CA, ID, MT, OR, SD
Diadesmis confervacea Kützing, 1844 Diadesmis contenta (Grunow ex Van Heurck) Mann in Round, Crawford & Mann, 1990
CA, MT CA, ID, MT, OR, WA, WY
Diadesmis contenta var. biceps (Grunow in Van Heurck) Hamilton in Hamilton, Poulin, Charles & Angell, 1992
CA, MT
Diadesmis gallica W. Smith, 1857 Diadesmis laevissima (Cleve) Mann in Round, Crawford & Mann, 1990 Diadesmis perpusilla (Grunow) Mann in Round, Crawford & Mann, 1990
CA, ID, MT, NV, SD, WA MT CA, ID, MT, OR, WA, WY
Diatoma ehrenbergii Kützing, 1844 Diatoma hiemale (Lyngbye) Heiberg, 1863 Diatoma mesodon Kützing, 1844 Diatoma moniliforme Kützing, 1833 Diatoma tenue Agardh, 1824 Diatoma vulgare Bory, 1831
MT CA, ID, MT, OR, SD, WA, WY CA, ID, MT, OR, SD, WA, WY CA, ID, MT, ND, OR, SD, WA, WY CA, ID, MT, ND, NV, OR, SD, WA, WY CA, ID, MT, NV, OR, SD, WA, WY
Diatomella balfouriana Greville, 1855
CA, ID, MT, OR, WA, WY
Dickieia subinflata (Grunow in Cleve & Möller) Mann, 1994
MT, WY
Didymosphenia geminata (Lyngbye) Schmidt in Schmidt et al., 1899
CA, ID, MT, ND, OR, WA, WY
Diploneis boldtiana Cleve, 1891 Diploneis elliptica (Kützing) Cleve, 1894 Diploneis finnica (Ehrenberg) Cleve, 1891
ID, MT, OR, WY CA, ID, MT, OR, SD, WA, WY ID, MT
A CHECKLIST OF DIATOMS FROM INLAND WATERS OF THE NORTHWESTERN U.S.
13
Checklist cont. Diploneis latefurcata (Fontell) Cleve-Euler, 1953 Diploneis marginestriata Hustedt, 1922 Diploneis modica Hustedt, 1945 Diploneis oblongella (Naegeli in Kützing) Cleve-Euler, 1922 Diploneis oculata (Brébisson in Desmazières) Cleve, 1894 Diploneis ovalis (Hilse in Rabenhorst) Cleve, 1891 Diploneis parma Cleve, 1891 Diploneis petersenii Hustedt, 1937 Diploneis pseudovalis Hustedt, 1930 Diploneis puella (Schumann) Cleve, 1894 Diploneis smithii (Brébisson in W. Smith) Cleve, 1894 Diploneis smithii var. pumila (Grunow) Hustedt, 1937 Diploneis subovalis Cleve, 1894
MT CA, ID, MT, OR, WY ID, MT CA, ID, MT, ND, OR, SD, WA, WY ID, MT, WY CA, ID, MT, ND, OR, WA, WY CA, ID, MT, SD, WY CA, MT, WA CA, ID, MT, SD, WY CA, ID, MT, ND, NV, OR, SD, WY CA, ID, MT, SD, WY MT CA, MT, OR, SD, WY
Discostella glomerata (Bachmann) Houk & Klee, 2004 Discostella pseudostelligera (Hustedt) Houk & Klee, 2004 Discostella stelligera (Cleve & Grunow) Houk & Klee, 2004 Discostella woltereckii (Hustedt) Houk & Klee, 2004
MT CA, ID, MT, OR, SD, WA, WY CA, ID, MT, OR, WA CA, WA
Distrionella incognita (Reichardt) Williams, 1990
ID, MT, ND, OR
Ellerbeckia arenaria (Moore ex Ralfs) Crawford, 1988
MT, OR, WY
Encyonema auerswaldii Rabenhorst, 1853 Encyonema brehmii (Hustedt) Mann in Round, Crawford & Mann, 1990 Encyonema brevicapitatum Krammer, 1997 Encyonema carina Lange-Bertalot & Krammer in Krammer, 1997 Encyonema cespitosum Kützing, 1849 Encyonema elginense (Krammer) Mann in Round, Crawford & Mann, 1990 Encyonema evergladianum Krammer, 1997 Encyonema fogedii Krammer, 1997 Encyonema gaeumannii (Meister) Krammer, 1997 Encyonema geisslerae Krammer, 1997 Encyonema hebridicum (Gregory) Grunow in Cleve & Möller, 1877 Encyonema hophense Krammer, 1997 Encyonema kamtschaticum Krammer, 1997 Encyonema lange-bertalotii Krammer, 1997 Encyonema lange-bertalotii var. obscuriformis Krammer, 1997 Encyonema latecapitatum Krammer, 1997 Encyonema latens (Krasske) Mann in Round, Crawford & Mann, 1990 Encyonema latum Krammer, 1997 Encyonema lunatum (W. Smith) Van Heurck, 1880 Encyonema minutiforme Krammer, 1997 Encyonema minutum (Hilse ex Rabenhorst) Mann in Round, Crawford & Mann, 1990 Encyonema muelleri (Hustedt) Mann in Round, Crawford & Mann, 1990 Encyonema neogracile Krammer, 1997 Encyonema neomesianum Krammer, 1997 Encyonema neomuelleri Krammer, 1997 Encyonema norvegica (Grunow in Schmidt et al.) Bukhtiyarova, 1995 Encyonema obscurum (Krasske) Mann in Round, Crawford & Mann, 1990
CA, ID, MT, OR, SD, WA, WY ID, MT CA, ID, MT, WY SD CA, ID, MT, OR, WA, WY MT MT, ND, SD, WY MT, OR, SD, WA, WY WA CA, MT MT, SD, WY CA, MT, WY ID, MT, OR, SD, WA CA, MT MT MT ID, MT, WY OR ID, MT, ND CA, MT, OR, WA, WY CA, ID, MT, NV, OR, SD, WA, WY CA, ID, MT, ND, NV, OR, SD, WA, WY CA, ID, MT, OR, WA, WY ID, MT, OR, WA SD ID, MT MT
14
L.L. BAHLS
Checklist cont. Encyonema pankowii Lange-Bertalot & Krammer in Krammer, 1997 Encyonema pergracile Krammer, 1997 Encyonema perminutum Krammer, 1997 Encyonema perpusillum (Cleve) Mann in Round, Crawford & Mann, 1990 Encyonema procerum Krammer, 1997 Encyonema prostratum Kützing, 1844 Encyonema reichardtii (Krammer) Mann in Round, Crawford & Mann, 1990 Encyonema semilanceolatum Krammer, 1997 Encyonema silesiacum (Bleisch ex Rabenhorst) Mann in Round, Crawford & Mann, 1990 Encyonema simile Krammer, 1997 Encyonema sinicum Krammer, 1997 Encyonema subelginense Krammer, 1997 Encyonema subminutum Krammer, 1997 Encyonema subnorvegicum Krammer, 1997 Encyonema subobscurum Krammer, 1997 Encyonema supergracile Krammer & Lange-Bertalot in Krammer, 1997 Encyonema tenuissimum (Hustedt) Mann in Round, Crawford & Mann, 1990 Encyonema triangulum (Ehrenberg) Kützing, 1849 Encyonema ventricosum (Kützing) Grunow in Schmidt et al., 1875 Encyonema ventricosum var. angusta Krammer, 1997 Encyonema vulgare Krammer, 1997 Encyonema yellowstonianum Krammer, 1997
MT MT ID, MT, OR, WA ID, MT, OR, WA MT, OR CA, ID, MT, OR, SD, WA, WY CA, ID, MT, NV, OR, SD, WA, WY CA, MT, SD CA, ID, MT, ND, NV, OR, SD, WA, WY CA, MT, OR, WA ID MT CA, MT, WA ID MT MT MT CA, ID, MT, ND, WY CA, ID, MT, OR, SD, WA, WY MT CA, ID, MT, OR, WY ID, MT, OR, WY
Encyonopsis aequalis (W. Smith) Krammer, 1997 Encyonopsis alpina Krammer & Lange-Bertalot in Krammer, 1997 Encyonopsis cesatiformis Krammer, 1997 Encyonopsis cesatii (Rabenhorst) Krammer, 1997 Encyonopsis delicatissima (Hustedt) Krammer, 1997 Encyonopsis descripta (Hustedt) Krammer, 1997 Encyonopsis eifelana Krammer, 1997 Encyonopsis falaisensis (Grunow) Krammer, 1997 Encyonopsis krammeri Reichardt, 1997 Encyonopsis microcephala (Grunow) Krammer, 1997 Encyonopsis microcephala var. robusta (Hustedt) Krammer, 1997 Encyonopsis minuta Krammer & Reichardt in Krammer, 1997 Encyonopsis moseri Krammer & Lange-Bertalot in Krammer, 1997 Encyonopsis perborealis Krammer, 1997 Encyonopsis radialis Krammer, 1997 Encyonopsis subminuta Krammer & Reichardt in Krammer, 1997 Encyonopsis subspicula Krammer, 1997 Encyonopsis thienemannii (Hustedt) Krammer, 1997
OR MT MT CA, ID, MT, OR, WA, WY WY MT, ND, SD, WA, WY SD MT, SD CA, MT, ND, OR, SD, WY CA, ID, MT, ND, OR, SD, WA, WY MT CA, ID, MT, WY MT MT MT CA, ID, MT, ND, OR, SD, WA, WY MT ID
Entomoneis alata Ehrenberg, 1845 Entomoneis ornata (Bailey) Reimer in Patrick & Reimer, 1975 Entomoneis paludosa (W. Smith) Reimer in Patrick & Reimer, 1975 Entomoneis punctulata (Grunow) Osada & Kobayasi, 1990
CA, MT, ND, SD, WY ID, MT, ND, WY CA, MT, ND, OR, SD, WY MT
Eolimna minima (Grunow in Van Heurck) Lange-Bertalot in Moser, Lange-Bertalot & Metzeltin, 1998
CA, ID, MT, ND, NV, OR, SD, WA, WY
A CHECKLIST OF DIATOMS FROM INLAND WATERS OF THE NORTHWESTERN U.S.
15
Checklist cont. Epithemia adnata (Kützing) Brébisson, 1838 Epithemia adnata var. porcellus (Kützing) Patrick in Patrick & Reimer, 1975 Epithemia argus (Ehrenberg) Kützing, 1844 Epithemia smithii Carruthers, 1864 Epithemia sorex Kützing, 1844 Epithemia turgida (Ehrenberg) Kützing, 1844 Epithemia turgida var. granulata (Ehrenberg) Brun, 1880 Epithemia turgida var. westermannii (Ehrenberg) Grunow, 1862
CA, ID, MT, ND, NV, OR, SD, WA, WY MT CA, MT, ND, OR MT CA, ID, MT, ND, NV, OR, SD, WA, WY CA, ID, MT, ND, NV, OR, SD, WA, WY MT MT
Eucocconeis alpestris (Brun) Lange-Bertalot in Lange-Bertalot & Genkal, 1999 Eucocconeis depressa (Cleve) Lange-Bertalot in Lange-Bertalot & Genkal, 1999 Eucocconeis flexella (Kützing) Cleve, 1895 Eucocconeis laevis (Østrup) Lange-Bertalot in Lange-Bertalot & Genkal, 1999 Eucocconeis lapponica Hustedt, 1924
MT MT CA, ID, MT, WA, WY CA, ID, MT, ND, OR, WA, WY MT
Eunotia arcubus Norpel & Lange-Bertalot in Lange-Bertalot, 1993 Eunotia arcus Ehrenberg, 1838 Eunotia bilunaris (Ehrenberg) Mills, 1934 Eunotia botuliformis Wild, Nörpel & Lange-Bertalot in Lange-Bertalot, 1993 Eunotia diadema Ehrenberg, 1837 Eunotia diodon Ehrenberg, 1837 Eunotia eruca Ehrenberg, 1854 Eunotia exigua (Brébisson ex Kützing) Rabenhorst, 1864 Eunotia faba Ehrenberg, 1838 Eunotia flexuosa (Brébisson in Kützing) Kützing, 1849 Eunotia formica Ehrenberg, 1843 Eunotia glacialis Meister, 1912 Eunotia implicata Nörpel, Lange-Bertalot & Alles in Alles, Nörpel & Lange-Bertalot, 1991 Eunotia incisa Gregory, 1854 Eunotia intermedia (Krasske ex Hustedt) Nörpel & Lange-Bertalot in Lange-Bertalot, 1993 Eunotia minor (Kützing) Grunow in Van Heurck, 1882-1885 Eunotia monodon Ehrenberg, 1843 Eunotia monodon var. bidens (Gregory) Hustedt, 1932 Eunotia muscicola Krasske, 1939 Eunotia muscicola var. tridentula Nörpel & Lange-Bertalot in Lange-Bertalot, 1993 Eunotia naegelii Migula, 1907 Eunotia nymanniana Grunow in Van Heurck, 1882-1885 Eunotia paludosa Grunow, 1862 Eunotia pectinalis (Kützing) Rabenhorst, 1864 Eunotia praerupta Ehrenberg, 1843 Eunotia praerupta var. bidens (Ehrenberg) Grunow in Cleve & Grunow, 1880 Eunotia rhomboidea Hustedt, 1950 Eunotia septentrionalis Østrup, 1897 Eunotia serpentina Ehrenberg, 1854 Eunotia siolii Hustedt, 1952 Eunotia soleirolii (Kützing) Rabenhorst, 1864 Eunotia steineckei Petersen, 1930 Eunotia subarcuatoides Alles, Nörpel & Lange-Bertalot, 1991 Eunotia tenella (Grunow in Van Heurck) Hustedt in Schmidt et al., 1913 Eunotia tetraodon Ehrenberg, 1838
OR CA, ID, MT, OR, WA CA, ID, MT, ND, OR, WA, WY MT MT ID, MT OR ID, MT, OR, SD, WA ID MT, OR CA, MT, OR, WA CA, MT, OR CA, ID, MT CA, ID, MT, OR, WA, WY MT CA, ID, MT, OR, SD, WA, WY ID, MT, OR, WA MT CA, ID, MT, OR, WA, WY CA, ID, MT CA, MT, OR MT CA, ID, MT, OR ID, MT, WA ID, MT, OR, WA MT ID, MT, OR, WA MT, WY CA, OR OR ID, MT, OR, WA MT CA, MT, OR, WA MT, OR, WA MT
16
L.L. BAHLS
Checklist cont. Eunotia triodon Ehrenberg, 1837 Eunotia valida Hustedt, 1930 Eunotia veneris (Kützing) De Toni, 1892
MT MT MT
Fallacia insociabilis (Krasske) Mann in Round, Crawford & Mann, 1990 Fallacia lenzii (Hustedt) Lange-Bertalot in Werum & Lange-Bertalot, 2004 Fallacia litoricola (Hustedt) Mann in Round, Crawford & Mann, 1990 Fallacia monoculata (Hustedt) Mann in Round, Crawford & Mann, 1990 Fallacia omissa (Hustedt) Mann in Round, Crawford & Mann, 1990 Fallacia pygmaea (Kützing) Stickle & Mann in Round, Crawford & Mann, 1990 Fallacia subhamulata (Grunow in Van Heurck) Mann in Round, Crawford & Mann, 1990 Fallacia tenera (Hustedt) Mann in Round, Crawford & Mann, 1990
MT CA, ID, MT, WA, WY CA CA, ID, MT, OR, SD, WY CA, ID, MT, ND, WY CA, ID, MT, ND, NV, OR, SD, WY CA, ID, MT, OR, WY CA, ID, MT, ND, SD, WY
Fistulifera pelliculosa (Brébisson) Lange-Bertalot, 1997
ID, MT, ND, NV, SD, WY
Fragilaria amphicephala Ehrenberg, 1854 Fragilaria berolinensis (Lemmermann) Lange-Bertalot, 1993 Fragilaria capucina Desmaziéres, 1825 Fragilaria capucina var. austriaca (Grunow) Lange-Bertalot, 1980 Fragilaria capucina var. capitellata (Grunow) Lange-Bertalot, 1991 Fragilaria capucina var. gracilis (Østrup) Hustedt, 1950 Fragilaria capucina var. mesolepta (Rabenhorst) Rabenhorst, 1864 Fragilaria crotonensis Kitton, 1869 Fragilaria neoproducta Lange-Bertalot in Krammer & Lange-Bertalot, 1991 Fragilaria nitzschioides Grunow in Van Heurck, 1881 Fragilaria sepes Ehrenberg, 1854 Fragilaria subsalina (Grunow) Lange-Bertalot in Krammer & Lange-Bertalot, 1991 Fragilaria tenera (W. Smith) Lange-Bertalot, 1980 Fragilaria vaucheriae (Kützing) Petersen, 1938 Fragilaria vaucheriae var. capitellata (Grunow) Patrick in Patrick & Freese, 1961 Fragilaria zeilleri Héribaud, 1903
MT CA CA, ID, MT, ND, NV, OR, SD, WA, WY MT MT CA, ID, MT, OR ID, MT, ND, OR, SD, WY CA, ID, MT, ND, NV, OR, SD, WY ID, MT CA, ID, MT, OR, WA, WY CA, ID, MT, ND, OR, SD, WY MT ID, MT, ND, SD, WY CA, ID, MT, ND, NV, OR, SD, WA, WY MT ID, MT
Fragilariforma bicapitata (Mayer) Williams & Round, 1988 Fragilariforma lata (Cleve-Euler) Williams & Round, 1988 Fragilariforma strangulata (Zanon) Williams & Round, 1988 Fragilariforma virescens (Ralfs) Williams & Round, 1988
CA, ID, MT, OR, WA, WY ID, MT MT CA, ID, MT, NV, OR, WA, WY
Frustulia amosseana Lange-Bertalot in Rumrich, Lange-Bertalot & Rumrich, 2000 Frustulia amphipleuroides (Grunow in Cleve & Grunow) Cleve-Euler, 1934 Frustulia crassinervia (Brébisson) Lange-Bertalot & Krammer in Lange-Bertalot & Metzeltin, 1996 Frustulia erifuga Lange-Bertalot & Krammer in Lange-Bertalot & Metzeltin, 1996 Frustulia krammeri Lange-Bertalot & Metzeltin in Metzeltin & Lange-Bertalot, 1998 Frustulia quadrisinuata Lange-Bertalot in Lange-Bertalot & Metzeltin, 1996 Frustulia rhomboides (Ehrenberg) DeToni, 1891 Frustulia saxonica Rabenhorst, 1848-1860 Frustulia spicula Amossé, 1932 Frustulia vulgaris (Thwaites) DeToni, 1891 Frustulia weinholdii Hustedt in Schmidt et al., 1936
CA CA, ID, MT, OR, WA, WY CA, ID, MT, WA, WY MT CA, MT MT CA, ID, MT, OR, WA, WY CA, ID, MT MT CA, ID, MT, ND, NV, OR, SD, WA, WY WA
Geissleria acceptata (Hustedt) Lange-Bertalot & Metzeltin, 1996
CA, ID, MT, NV, OR, SD, WA, WY
A CHECKLIST OF DIATOMS FROM INLAND WATERS OF THE NORTHWESTERN U.S.
17
Checklist cont. Geissleria cummerowi (Kalbe) Lange-Bertalot, 2001 Geissleria decussis (Østrup) Lange-Bertalot & Metzeltin, 1996 Geissleria dolomitica (Bock) Lange-Bertalot & Metzeltin, 1996 Geissleria ignota (Krasske) Lange-Bertalot & Metzeltin, 1996 Geissleria paludosa (Hustedt) Lange-Bertalot & Metzeltin, 1996 Geissleria punctifera (Hustedt) Metzeltin, Lange-Bertalot & García-Rodríguez, 2005 Geissleria schoenfeldii (Hustedt) Lange-Bertalot & Metzeltin, 1996 Geissleria similis (Krasske) Lange-Bertalot & Metzeltin, 1996
OR CA, ID, MT, NV, OR, SD, WA, WY MT CA, ID, MT, OR, SD, WA, WY CA, MT, OR, WA, WY CA ID, MT, OR, WY MT
Germainiella enigmatica (Germain) Metzeltin, Lange-Bertalot & García-Rodríguez, 2005
CA
Gomphoneis eriense (Grunow) Skvortzow & Meyer, 1928 Gomphoneis eriense var. variabilis Kociolek & Stoermer, 1988 Gomphoneis herculeanum (Ehrenberg) Cleve, 1894 Gomphoneis linearis Kociolek & Stoermer, 1986 Gomphoneis mammilla (Ehrenberg) Cleve, 1894 Gomphoneis minuta (Stone) Kociolek & Stoermer, 1988 Gomphoneis septa (Moghadam) Kociolek, Stoermer & Bahls, 1986 Gomphoneis trullata Kociolek & Stoermer, 1986
CA, ID, MT, ND, NV, OR, SD, WA, WY MT CA, ID, MT, WA, WY ID, MT, OR OR CA, ID, MT, OR, WA, WY ID, MT, WY OR
Gomphonema acidoclinatum Lange-Bertalot & Reichardt in Werum & Lange-Bertalot, 2004 Gomphonema acuminatum Ehrenberg, 1832 Gomphonema affine Kützing, 1844 Gomphonema americobtusatum Reichardt & Lange-Bertalot in Reichardt, 1999 Gomphonema amoenum Lange-Bertalot in Krammer & Lange-Bertalot, 1985 Gomphonema anglicum Ehrenberg, 1843 Gomphonema angustatum (Kützing) Rabenhorst, 1864 Gomphonema angusticephalum Reichardt & Lange-Bertalot in Reichardt, 1999 Gomphonema angustivalva Reichardt, 1997 Gomphonema angustum Agardh, 1831 Gomphonema anjae Lange-Bertalot & Reichardt, 1991 Gomphonema apuncto Wallace, 1960 Gomphonema aquaemineralis Lange-Bertalot & Reichardt, 1991 Gomphonema augur Ehrenberg, 1840 Gomphonema auritum Braun ex Kützing, 1849 Gomphonema bipunctatum Krasske, 1943 Gomphonema bohemicum Reichelt & Fricke in Schmidt et al., 1902 Gomphonema bourbonense Reichardt, 1997 Gomphonema bozenae Lange-Bertalot & Reichardt in Lange-Bertalot & Metzeltin, 1996 Gomphonema brebissonii Kützing, 1849 Gomphonema capitatum Ehrenberg, 1838 Gomphonema citera Hohn & Hellerman, 1963 Gomphonema clava Reichardt, 2001 Gomphonema clavatulum Reichardt, 1999 Gomphonema clavatum Ehrenberg, 1838 Gomphonema clevei Fricke in Schmidt et al., 1902 Gomphonema consector Hohn & Hellerman, 1963 Gomphonema contraturris Lange-Bertalot & Reichardt in Lange-Bertalot, 1993 Gomphonema coronatum Ehrenberg, 1840 Gomphonema cuneolus Reichardt, 1997
MT CA, ID, MT, NV, OR, WA, WY CA, ID, MT, ND, SD, WA, WY CA, ID, MT, OR, WA, WY ID, MT, OR, WY MT CA, ID, MT, ND, NV, OR, SD, WA, WY MT ID, MT, OR, WA, WY CA, ID, MT, OR, SD, WA, WY MT MT ID, MT, WY ID, MT, ND, WY CA, MT, OR, WA CA, MT, OR, WA, WY MT, WA ID ID, MT CA, ID, MT, ND, SD, WA, WY MT, ND, OR, WY CA, MT, WY MT MT, OR, WY CA, ID, MT, ND, NV, OR, SD, WA, WY CA, ID, MT, NV, OR, WA, WY MT MT MT ID, MT, OR
18
L.L. BAHLS
Checklist cont. Gomphonema cymbelliclinum Reichardt & Lange-Bertalot in Reichardt, 1999 Gomphonema designatum Reichardt, 1997 Gomphonema dichotomum Kützing, 1833 Gomphonema drutelingense Reichardt, 1999 Gomphonema entolejum Østrup, 1902
Gomphonema exilissimum (Grunow) Lange-Bertalot & Reichardt in Lange-Bertalot & Metzeltin, 1996
Gomphonema extentum Reichardt & Lange-Bertalot in Reichardt, 1994 Gomphonema geisslerae Reichardt & Lange-Bertalot in Reichardt, 1997 Gomphonema geitleri Kociolek & Stoermer, 1991 Gomphonema gracile Ehrenberg, 1838 Gomphonema grasmueckii Lange-Bertalot & Reichardt in Lange-Bertalot, 1993 Gomphonema hebridense Gregory, 1854 Gomphonema hedinii Hustedt, 1922 Gomphonema innocens Reichardt, 1999 Gomphonema insigne Gregory, 1856 Gomphonema insigniforme Reichardt & Lange-Bertalot in Reichardt, 1999 Gomphonema instabilis Hohn & Hellerman, 1963 Gomphonema interpositum Reichardt, 1999 Gomphonema intricatum Kützing, 1844 Gomphonema intricatum var. vibrio (Ehrenberg) Cleve, 1894 Gomphonema kobayasii Kociolek & Kingston, 1999 Gomphonema lagenula Kützing, 1844 Gomphonema lateripunctatum Reichardt & Lange-Bertalot, 1991 Gomphonema laticollum Reichardt, 2001 Gomphonema longilineare Reichardt, 1999 Gomphonema maclaughlinii Reichardt, 1999 Gomphonema manubrium Fricke in Schmidt et al., 1904 Gomphonema mehleri Camburn, 1978 Gomphonema mexicanum Grunow in Van Heurck, 1880 Gomphonema micropumilum Reichardt, 1997 Gomphonema micropus Kützing, 1844 Gomphonema minuscula Krasske, 1932 Gomphonema minutiforme Lange-Bertalot & Reichardt in Krammer & Lange-Bertalot, 1991 Gomphonema minutum Agardh, 1831 Gomphonema montanum Schumann, 1867 Gomphonema occultum Reichardt & Lange-Bertalot, 1991 Gomphonema olivaceoides Hustedt, 1950 Gomphonema olivaceum (Hornemann) Brébisson, 1838 Gomphonema olivaceum var. calcareum (Cleve) Van Heurck, 1880 Gomphonema pala Reichardt, 2001 Gomphonema paludosum Reichardt, 1999 Gomphonema parallelistriatum Lange-Bertalot & Reichardt in Lange-Bertalot, 1993
Gomphonema parvulius (Lange-Bertalot & Reichardt) Lange-Bertalot & Reichardt in LangeBertalot & Metzeltin, 1996
Gomphonema parvulum (Kützing) Kützing, 1849 Gomphonema procerum Reichardt & Lange-Bertalot, 1991 Gomphonema productum (Grunow) Lange-Bertalot & Reichardt in Lange-Bertalot, 1993 Gomphonema pseudoaugur Lange-Bertalot, 1979 Gomphonema pseudobohemicum Lange-Bertalot & Reichardt in Lange-Bertalot & Metzeltin, 1996
CA, ID, MT, OR, WY CA, MT, ND, OR CA, ID, MT, OR MT, SD, WY CA CA, MT, WY MT MT MT CA, ID, MT, ND, OR, SD, WA, WY MT, OR MT, OR CA, MT, OR, WY CA, ID, MT, OR, SD, WA, WY CA, ID, MT, WA, WY CA, ID, MT, OR MT MT CA, ID, MT, OR, SD CA, MT, OR CA, ID, MT, ND, OR, SD, WA, WY CA, MT, WY CA, ID, MT, WY CA, MT, ND CA, ID, MT, ND, SD, WA, WY CA, MT, OR, SD, WY MT MT CA, ID, MT, ND, OR, SD, WA, WY CA, ID, MT, OR, WA, WY CA, ID, MT, ND, NV, OR, SD, WA, WY CA, ID, MT, OR, WA, WY CA, ID, MT, OR, WY CA, ID, MT, ND, NV, OR, SD, WA, WY MT, WY MT CA, ID, MT, OR, WA, WY CA, ID, MT, ND, NV, OR, SD, WA, WY MT CA, ID, MT, ND MT, OR MT CA, ID, MT, WY CA, ID, MT, ND, NV, OR, SD, WA, WY CA, ID, MT, OR, WA, WY ID, MT, OR MT CA, MT, SD
A CHECKLIST OF DIATOMS FROM INLAND WATERS OF THE NORTHWESTERN U.S.
19
Checklist cont. Gomphonema pseudopusillum Reichardt, 1999 Gomphonema pseudotenellum Lange-Bertalot in Krammer & Lange-Bertalot, 1985 Gomphonema pumilum (Grunow) Reichardt & Lange-Bertalot, 1991 Gomphonema pygmaeum Kociolek & Stoermer, 1991 Gomphonema quadripunctatum (Østrup) Wislouch, 1924 Gomphonema reimeri Kociolek & Kingston, 1999 Gomphonema rhombicum Fricke in Schmidt et al., 1904 Gomphonema sarcophagus Gregory, 1856 Gomphonema semiapertum Grunow in Van Heurck, 1880 Gomphonema sinestigma Reichardt, Jüttner & Cox in Jüttner et al., 2004 Gomphonema sphaerophorum Ehrenberg, 1845 Gomphonema stoermeri Kociolek & Kingston, 1999 Gomphonema stonei Reichardt, 1999 Gomphonema subclavatum (Grunow in Van Heurck) Grunow, 1884 Gomphonema subtile Ehrenberg, 1843 Gomphonema tergestinum (Grunow in Van Heurck) Fricke in Schmidt et al., 1902 Gomphonema truncatum Ehrenberg, 1832 Gomphonema utae Lange-Bertalot & Reichardt in Reichardt, 1999 Gomphonema ventricosum Gregory, 1856 Gomphonema vibrio Ehrenberg, 1843 Gomphonema vibrioides Reichardt & Lange-Bertalot, 1991
MT CA, MT, OR, WY CA, ID, MT, ND, NV, OR, SD, WA, WY MT MT MT CA, ID, MT, NV, OR, WA, WY ID, MT, ND, OR, SD, WY OR MT CA, ID, MT OR SD CA, ID, MT, ND, OR, SD, WA, WY ID, MT, ND MT CA, ID, MT, ND, NV, OR, SD, WA, WY ID, MT, WY WA CA, MT, OR ND
Gomphosphenia tackei (Hustedt) Lange-Bertalot, 1995
CA, ID, MT, OR
Gyrosigma acuminatum (Kützing) Rabenhorst, 1853 Gyrosigma attenuatum (Kützing) Rabenhorst, 1853 Gyrosigma balticum (Ehrenberg) Rabenhorst, 1853 Gyrosigma macrum (W. Smith) Griffith & Henfrey, 1856 Gyrosigma nodiferum (Grunow) G. West, 1907 Gyrosigma obtusatum (Sullivant & Wormley) Boyer, 1922 Gyrosigma peisonis (Grunow) Hustedt, 1930 Gyrosigma scalproides (Rabenhorst) Cleve, 1894 Gyrosigma sciotense (Sullivant & Wormley) Cleve, 1894 Gyrosigma wansbeckii (Donkin) Cleve, 1894 Gyrosigma wormleyi (Sullivant in Sullivant & Wormley) Boyer, 1922
CA, ID, MT, ND, NV, OR, SD, WA, WY CA, ID, MT, OR, SD, WY MT MT, SD, WY CA, ID, MT, WY CA, ID, MT MT, SD CA, ID, MT MT, SD WY MT
Hannaea arcus (Ehrenberg) Patrick in Patrick & Freese, 1961 Hannaea arcus var. amphioxys (Rabenhorst) Patrick in Patrick & Reimer, 1966
CA, ID, MT, ND, NV, OR, WA, WY MT
Hantzschia abundans Lange-Bertalot, 1993 Hantzschia amphioxys (Ehrenberg) Grunow in Cleve & Grunow, 1880 Hantzschia distinctepunctata (Hustedt) Hustedt in Schmidt et al., 1921 Hantzschia elongata (Hantzsch) Grunow in Cleve & Grunow, 1880 Hantzschia rhaetica Meister, 1912 Hantzschia spectabilis (Ehrenberg) Hustedt, 1959 Hantzschia subrupestris Lange-Bertalot, 1993 Hantzschia virgata (Roper) Grunow in Cleve & Grunow, 1880 Hantzschia vivacior Lange-Bertalot, 1993 Hantzschia vivax (W. Smith) Peragallo in Tempère & Peragallo, 1908
CA CA, ID, MT, ND, NV, OR, SD, WA, WY MT MT MT MT MT CA, MT, SD, WY CA MT
20
L.L. BAHLS
Checklist cont. Haslea spicula (Hickie) Lange-Bertalot, 1997
WY
Hippodonta avittata (Cholnoky) Lange-Bertalot, Metzeltin & Witkowski, 1996 Hippodonta capitata (Ehrenberg) Lange-Bertalot, Metzeltin & Witkowski, 1996 Hippodonta coxiae Lange-Bertalot, 2001 Hippodonta hungarica (Grunow) Lange-Bertalot, Metzeltin & Witkowski, 1996 Hippodonta lueneburgensis (Grunow) Lange-Bertalot, Metzeltin & Witkowski, 1996 Hippodonta umbilicatissima (Reichardt) Reichardt in Lange-Bertalot, Metzeltin & Witkowski, 1996
MT CA, ID, MT, ND, NV, OR, SD, WA, WY CA CA, ID, MT, ND, OR, SD, WY ID, MT, WA MT
Hydrosera triquetra Wallich, 1858
CA
Karayevia amoena (Hustedt) Bukhtiyarova, 2006 Karayevia clevei (Grunow in Cleve & Grunow) Round & Bukhtiyarova, 1996 Karayevia laterostrata (Hustedt) Round & Bukhtiyarova, 1996 Karayevia ploenensis (Hustedt) Bukhtiyarova, 2006 Karayevia suchlandtii (Hustedt) Bukhtiyarova, 2006
ID CA, ID, MT, ND, NV, OR, WA, WY CA, ID, MT, OR, SD, WA, WY ID, MT CA, ID, MT, OR, WA, WY
Kobayasiella subtilissima (Cleve) Lange-Bertalot, 1999
MT, WA
Lemnicola hungarica (Grunow) Round & Basson, 1997
CA, ID, MT, ND, OR, WA, WY
Luticola acidoclinata Lange-Bertalot in Lange-Bertalot & Metzeltin, 1996 Luticola cohnii (Hilse) Mann in Round, Crawford & Mann, 1990 Luticola goeppertiana (Bleisch in Rabenhorst) Mann in Round, Crawford & Mann, 1990 Luticola mutica (Kützing) Mann in Round, Crawford & Mann, 1990 Luticola muticopsis (Van Heurck) Mann in Round, Crawford & Mann, 1990 Luticola nivalis (Ehrenberg) Mann in Round, Crawford & Mann, 1990 Luticola obligata (Hustedt) Mann in Round, Crawford & Mann, 2000 Luticola saxophila (Bock ex Hustedt) Mann in Round, Crawford & Mann, 1990 Luticola undulata (Hilse ex Rabenhorst) Mann in Round, Crawford & Mann, 1990
MT ID, MT, WY CA, ID, MT, OR, WA CA, ID, MT, ND, NV, OR, SD, WA, WY CA, ID, MT, ND, OR CA, ID, MT, NV, OR CA CA MT
Mastogloia elliptica (Agardh) Cleve in Schmidt et al., 1893 Mastogloia elliptica var. dansei (Thwaites) Cleve, 1895 Mastogloia grevillei W. Smith in Gregory, 1856 Mastogloia pumila (Grunow in Van Heurck) Cleve, 1895 Mastogloia smithii Thwaites ex W. Smith, 1856 Mastogloia smithii var. lacustris Grunow, 1878
MT, ND, SD, WY MT MT, ND, WY WY MT, ND, SD, WY MT
Mayamaea agrestis (Hustedt) Lange-Bertalot, 2001 Mayamaea aliena (Krasske) Lange-Bertalot, 2001 Mayamaea atomus (Kützing) Lange-Bertalot, 1997 Mayamaea atomus var. permitis (Hustedt) Lange-Bertalot, 1997 Mayamaea excelsa (Krasske) Lange-Bertalot, 1997 Mayamaea fossalis (Krasske) Lange-Bertalot, 1997
CA, ID, MT, OR, SD, WA, WY MT CA, ID, MT, ND, OR, SD, WA, WY CA, ID, MT, NV, OR, WA, WY MT ID, MT, OR
Melosira varians Agardh, 1827
CA, ID, MT, ND, NV, OR, SD, WA, WY
Meridion anceps (Ehrenberg) Williams, 1985 Meridion circulare (Greville) Agardh, 1831 Meridion circulare var. constrictum (Ralfs) Van Heurck, 1881
MT, WA CA, ID, MT, OR, SD, WA, WY ID, MT, WY
A CHECKLIST OF DIATOMS FROM INLAND WATERS OF THE NORTHWESTERN U.S.
21
Checklist cont. Microcostatus krasskei (Hustedt) Johansen & Sray, 1998 Microcostatus kuelbsii (Lange-Bertalot in Krammer & Lange-Bertalot) Lange-Bertalot, 1999 Microcostatus maceria (Schimanski) Lange-Bertalot, 1999
WY MT MT, OR
Muelleria gibbula (Cleve) Spaulding & Stoermer, 1997
OR
Navicula absoluta Hustedt, 1950 Navicula ammophila Grunow, 1882 Navicula amphibola Cleve, 1891 Navicula amphiceropsis Lange-Bertalot & Rumrich in Rumrich, Lange-Bertalot & Rumrich, 2000 Navicula angusta Grunow, 1860 Navicula antonii Lange-Bertalot & Rumrich in Rumrich, Lange-Bertalot & Rumrich, 2000 Navicula aquaedurae Lange-Bertalot, 1993
CA, ID, MT, OR MT, ND, WY MT CA, MT, SD, WY CA, ID, MT, OR CA, ID, MT, ND, NV, OR, SD, WA, WY CA, ID, MT, OR, WA, WY
Navicula arctotenelloides Lange-Bertalot & Metzeltin in Lange-Bertalot, Kulbs, Lauser, NorpelSchempp & Willmann, 1996
Navicula arenaria Donkin, 1861 Navicula arkona Lange-Bertalot & Witkowski, 2001 Navicula arvensis Hustedt, 1937 Navicula asellus Weinhold ex Hustedt in Schmidt et al., 1934 Navicula associata Lange-Bertalot, 2001 Navicula aurora Sovereign, 1958 Navicula austrocollegarum Lange-Bertalot & Voigt, 2001 Navicula bacilloides Hustedt, 1945 Navicula bicephala Hustedt, 1952 Navicula bjoernoeyaensis Metzeltin, Witkowski & Lange-Bertalot in Metzeltin & Witkowski, 1996 Navicula bourrellyivera Lange-Bertalot, Witkowski & Stachura in Witkowski, Lange-Bertalot & Stachura, 1998
Navicula brekkaensis Petersen, 1928 Navicula brockmannii Hustedt, 1934 Navicula broetzii Lange-Bertalot & Reichardt in Lange-Bertalot & Metzeltin, 1996 Navicula canalis Patrick, 1944 Navicula capitatoradiata Germain, 1981 Navicula cari Ehrenberg, 1836 Navicula cariocincta Lange-Bertalot, 1993 Navicula cascadensis Sovereign, 1958 Navicula catalanogermanica Lange-Bertalot & Hofmann in Lange-Bertalot, 1993 Navicula cataracta-rheni Lange-Bertalot, 1993 Navicula caterva Hohn & Hellerman, 1963 Navicula chiarae Lange-Bertalot & Genkal, 1999 Navicula cincta (Ehrenberg) Ralfs in Pritchard, 1861 Navicula cincta var. rostrata Reimer, 1961 Navicula concentrica Carter & Bailey-Watts, 1981 Navicula constans Hustedt, 1944 Navicula convergens Patrick, 1959 Navicula cryptocephala Kützing, 1844 Navicula cryptocephaloides Hustedt, 1937 Navicula cryptofallax Lange-Bertalot & Hofmann in Lange-Bertalot, 1993 Navicula cryptotenella Lange-Bertalot in Krammer & Lange-Bertalot, 1985 Navicula cryptotenelloides Lange-Bertalot, 1993 Navicula curtisterna Lange-Bertalot, 2001
CA, MT, WY WY ID, MT, OR, WA CA, ID, MT, OR, SD, WA, WY CA, OR ID, MT, WY CA, ID, MT, OR, WA MT CA, ID, MT, NV, OR, SD, WY MT CA, MT, WY MT, WY MT ID, MT, OR MT, OR CA, MT, ND, OR, WY CA, ID, MT, ND, NV, OR, SD, WA, WY CA, ID, MT, ND, OR, WA, WY MT CA, MT, OR CA, ID, MT, OR, WA, WY CA, ID, MT, OR, WY CA, ID, MT, ND, NV, OR, SD, WY OR CA, ID, MT, ND, NV, OR, SD, WY ID, MT, ND, WY CA, ID, MT, OR, WY CA, ID, MT, OR, WY MT CA, ID, MT, ND, NV, OR, SD, WA, WY MT, SD MT, SD, WY CA, ID, MT, ND, NV, OR, SD, WA, WY CA, ID, MT, OR, SD, WY SD
22
L.L. BAHLS
Checklist cont. Navicula dealpina Lange-Bertalot, 1993 Navicula declivis Hustedt, 1934 Navicula densilineolata (Lange-Bertalot) Lange-Bertalot, 1993 Navicula detenta Hustedt, 1943 Navicula difficillima Hustedt, 1950 Navicula difficillimoides Hustedt, 1957 Navicula digitulus Hustedt, 1943 Navicula duerrenbergiana Hustedt in Schmidt et al., 1934 Navicula egregia Hustedt, 1942 Navicula eidrigiana Carter, 1979
MT, OR MT CA, ID, MT, OR, SD, WY ID, MT, NV, OR, WA, WY CA, MT, ND, OR, WA MT, WA ID, MT, WY MT, SD, WY MT CA, MT, ND, SD, WY
Navicula erifuga Lange-Bertalot in Krammer & Lange-Bertalot, 1985 Navicula exilis Kützing, 1844 Navicula farta Hustedt in Schmidt et al., 1934 Navicula fennica Hustedt, 1962 Navicula festiva Krasske, 1925 Navicula flanatica Grunow, 1860 Navicula fracta Hustedt, 1961 Navicula gerloffii Schimanski, 1978 Navicula germainii Wallace, 1960 Navicula glomus Carter & Bailey-Watts, 1981 Navicula goersii Bahls, 1983 Navicula gondwana Lange-Bertalot, 1993 Navicula gottlandica Grunow in Van Heurck, 1857 Navicula gregaria Donkin, 1861 Navicula hambergii Hustedt, 1924 Navicula harderi Hustedt in Brendemuhl, 1949 Navicula helensis Schulz, 1926 Navicula hintzii Lange-Bertalot, 1993 Navicula hofmanniae Lange-Bertalot, 1993 Navicula impexa Hustedt, 1961 Navicula indifferens Hustedt, 1942 Navicula ingenua Hustedt, 1957 Navicula ingrata Krasske, 1938 Navicula integra (W. Smith) Ralfs in Pritchard, 1861 Navicula jaagii Meister, 1935 Navicula joubaudii Germain, 1982 Navicula kohlmaieri Lange-Bertalot in Lange-Bertalot & Genkal, 1999 Navicula kotschyi Grunow, 1860 Navicula krammerae Lange-Bertalot in Lange-Bertalot & Metzeltin, 1996 Navicula lanceolata (Agardh) Ehrenberg, 1838 Navicula laterostrata Hustedt, 1925 Navicula leistikowii Lange-Bertalot, 1993 Navicula leptostriata Jørgensen, 1948 Navicula libonensis Schoemann, 1970 Navicula ludloviana Schmidt in Schmidt et al., 1876 Navicula lundii Reichardt, 1985 Navicula lundstromii Cleve in Cleve & Grunow, 1880 Navicula margalithii Lange-Bertalot in Krammer & Lange-Bertalot, 1985
CA, ID, MT, ND, NV, OR, WD, WY CA, ID, MT, OR, SD, WA, WY MT ID OR MT MT ID, MT CA, ID, MT, OR, WA, WY MT CA, MT, ND, OR, SD, WY MT MT CA, ID, MT, ND, NV, OR, SD, WA, WY CA, ID, MT, OR CA, MT, OR MT CA, ID, MT, OR, SD, WY MT, WY OR CA, MT CA MT MT, ND, WY MT MT WY CA, ID, MT, SD, WY MT, OR, WY CA, ID, MT, NV, OR, SD, WA, WY MT CA, ID, MT, OR, SD, WY ID, MT, OR, WA CA, ID, MT, ND, NV, OR, SD, WA, WY OR CA, ID, MT, OR, WA, WY MT MT, SD, WY
Navicula elginensis var. cuneata (Möller ex Foged) Lange-Bertalot in Krammer & Lange-Bertalot, MT, OR 1985
A CHECKLIST OF DIATOMS FROM INLAND WATERS OF THE NORTHWESTERN U.S.
23
Checklist cont. Navicula medioconvexa Hustedt, 1961 Navicula mediocostata Reichardt, 1988 Navicula menisculus Schumann, 1867 Navicula meniscus Schumann, 1867 Navicula microcari Lange-Bertalot, 1993 Navicula microdigitoradiata Lange-Bertalot, 1993 Navicula minuta (Cleve) Cleve-Euler, 1953 Navicula modica Hustedt, 1945 Navicula moenofranconica Lange-Bertalot, 1993 Navicula moskalii Metzeltin, Witkowski & Lange-Bertalot in Metzeltin & Witkowski, 1996 Navicula muraloides Hustedt, 1950 Navicula namibica Lange-Bertalot & Rumrich in Lange-Bertalot, 1993 Navicula naumannii Hustedt, 1942 Navicula nipponica (Skvortzow) Lange-Bertalot, 1993 Navicula nolensoides Bock, 1970 Navicula normaloides Cholnoky, 1968 Navicula notha Wallace, 1960 Navicula novaesiberica Lange-Bertalot, 1993 Navicula nugalis Hohn & Hellerman, 1963 Navicula obdurata Hohn & Hellerman, 1963 Navicula oblonga (Kützing) Kützing, 1844 Navicula obsoleta Hustedt, 1942 Navicula occulta Krasske, 1929 Navicula ochridana Hustedt, 1945 Navicula oligotraphenta Lange-Bertalot & Hofmann in Lange-Bertalot, 1993 Navicula oppugnata Hustedt, 1945 Navicula ordinaria Hustedt, 1950 Navicula parablis Hohn & Hellerman, 1963 Navicula paracari Lange-Bertalot, 1993 Navicula paramutica Bock, 1963 Navicula pavillardi Hustedt, 1939 Navicula pennsylvanica Patrick in Patrick & Reimer, 1966 Navicula peregrina (Ehrenberg) Kützing, 1844 Navicula perminuta Grunow in Van Heurck, 1880 Navicula phyllepta Kützing, 1844 Navicula phylleptosoma Lange-Bertalot in Lange-Bertalot & Genkal, 1999 Navicula platystoma Ehrenberg, 1838 Navicula plausibilis Hustedt, 1964 Navicula porifera Hustedt, 1944 Navicula porifera var. opportuna (Hustedt) Lange-Bertalot in Krammer & Lange-Bertalot, 1985 Navicula praeterita Hustedt, 1945 Navicula pseudoarvensis Hustedt, 1942 Navicula pseudobryophila Hustedt, 1942 Navicula pseudokotschyi Lange-Bertalot in Krammer & Lange-Bertalot, 1985 Navicula pseudolanceolata Lange-Bertalot, 1980 Navicula pseudoppugnata Lange-Bertalot & Miho, 2001 Navicula pseudosilicula var. olympica Sovereign, 1963 Navicula pseudotenelloides Krasske, 1938 Navicula pseudoventralis Hustedt in Schmidt et al., 1936 Navicula pusio Cleve, 1895
CA, ID, MT, WY ID, MT, WY CA, ID, MT, ND, NV, OR, SD, WA, WY MT CA, ID, MT, WA MT MT MT CA, MT, WY ID, MT, WY MT CA, MT, ND, SD, WY SD MT WA CA, MT CA, ID, MT, ND, OR, SD, WY ID, MT, WY WA ID, MT, WA MT, ND, NV, OR, SD CA, MT, WY ID, MT MT CA, ID, MT, ND, OR, SD, WY CA, ID, MT, SD MT CA, MT CA, MT, OR ID, MT MT MT MT, ND, SD, WY CA, MT, ND, OR, WA, WY CA, ID, MT, ND, SD, WY CA, MT, SD MT ID, MT, OR MT, OR, WY MT MT CA, ID, MT, OR CA, ID, MT, WY WA CA, ID, MT, NV, OR, WA, WY CA, MT, WA ID, MT CA, ID, MT, ND, WY CA, ID, MT, OR ID, MT
24
L.L. BAHLS
Checklist cont. Navicula radiosa Kützing, 1844 Navicula radiosafallax Lange-Bertalot, 1993 Navicula radiosiola Lange-Bertalot, 1993 Navicula recens (Lange-Bertalot) Lange-Bertalot in Krammer & Lange-Bertalot, 1985 Navicula reichardtiana Lange-Bertalot in Lange-Bertalot & Krammer, 1989 Navicula reinhardtii (Grunow) Grunow, 1880 Navicula rhynchocephala Kützing, 1844 Navicula rhynchotella Lange-Bertalot, 1993 Navicula riediana Lange-Bertalot & Rumrich in Rumrich, Lange-Bertalot & Rumrich, 2000 Navicula rostellata Kützing, 1844 Navicula rotunda Hustedt, 1945 Navicula ruttneri var. rostrata Hustedt, 1937 Navicula salinarum Grunow, 1880 Navicula salinicola Hustedt, 1939 Navicula scabellum Hustedt, 1942 Navicula schadei Krasske, 1929 Navicula schmassmannii Hustedt, 1943 Navicula schroeteri Meister, 1932 Navicula seibigiana Lange-Bertalot, 1993 Navicula semen Ehrenberg, 1843 Navicula seminuloides Hustedt in Schmidt et al., 1936 Navicula similis Krasske, 1929 Navicula slesvicensis Grunow in Van Heurck, 1880 Navicula stankovici Hustedt, 1945 Navicula streckerae Lange-Bertalot & Witkowski in Witkowski, Lange-Bertalot & Metzeltin, 2000 Navicula stroemii Hustedt, 1931 Navicula subadnata Hustedt, 1942 Navicula subalpina Reichardt, 1988 Navicula subarvensis Hustedt, 1942 Navicula subconcentrica Lange-Bertalot, 2001 Navicula sublucidula Hustedt, 1950 Navicula subminuscula Manguin, 1942 Navicula submitis Hustedt, 1945 Navicula submuralis Hustedt, 1945 Navicula subnympharum Hustedt in Schmidt et al., 1936 Navicula subplacentula Hustedt in Schmidt et al., 1930 Navicula subrhynchocephala Hustedt, 1935 Navicula subrotundata Hustedt, 1945 Navicula suecorum Carlson, 1913 Navicula supergregaria Lange-Bertalot & Rumrich in Rumrich, Lange-Bertalot & Rumrich, 2000 Navicula symmetrica Patrick, 1944 Navicula tantula Hustedt, 1943 Navicula tenelloides Hustedt, 1937 Navicula tenerrima Hustedt, 1937 Navicula tridentula Krasske, 1923 Navicula tripunctata (O. Müller) Bory, 1822-1831 Navicula trivialis Lange-Bertalot, 1980 Navicula trophicatrix Lange-Bertalot in Lange-Bertalot & Metzeltin, 1996 Navicula upsaliensis (Grunow in Van Heurck) M. Peragallo, 1903 Navicula utermohli Hustedt, 1943
CA, ID, MT, ND, OR, SD, WA, WY ID, MT, OR, SD MT, OR CA, ID, MT, ND, OR, SD, WY CA, ID, MT, ND, NV, OR, SD, WA, WY ID, MT, NV, WA CA, ID, MT, ND, NV, OR, SD, WA, WY MT CA CA, ID, MT, ND, NV, OR, WY CA, MT, OR MT CA, MT, ND, SD, WY CA, ID, MT, ND, OR, SD, WA, WY OR MT ID, MT, WA, WY CA, MT, WA CA, MT, WA, WY CA, ID, MT ID, MT OR MT, ND, WY MT MT, ND, SD, WY CA, ID, MT, OR, SD, WA, WY ID, MT CA, MT, WY CA, MT MT MT, ND CA, ID, MT, ND, NV, OR, SD, WA, WY MT CA, ID, MT, WY ID ND, OR CA, MT, WY CA, ID, MT, OR MT MT CA, ID, MT, OR, SD, WA, WY ID, MT, WA CA, ID, MT, ND, NV, SD, WA, WY MT ID, MT, WY CA, ID, MT, ND, NV, OR, SD, WA, WY CA, ID, MT, ND, NV, OR, SD, WA, WY ID, MT, OR, WY CA, MT, OR, WY OR
A CHECKLIST OF DIATOMS FROM INLAND WATERS OF THE NORTHWESTERN U.S.
25
Checklist cont. Navicula vandamii Schoeman & Archibald, 1987 Navicula vaneei Lange-Bertalot in Witkowski, Lange-Bertalot & Stachura, 1998 Navicula variostriata Krasske, 1923 Navicula vaucheriae Petersen, 1915 Navicula venerablis Hohn & Hellerman, 1963 Navicula veneta Kützing, 1844 Navicula ventralis Krasske, 1923
Navicula vilaplanii (Lange-Bertalot & Sabater) Lange-Bertalot & Sabater in Rumrich, LangeBertalot & Rumrich, 2000
ID, MT, SD, WY CA, MT, WY CA, ID, MT, WA CA, MT, WA, WY CA, MT CA, ID, MT, ND, NV, OR, SD, WA, WY CA, ID, MT, OR, WA, WY CA, MT, SD, WY
Navicula viridula (Kützing) Ehrenberg, 1836 Navicula viridulacalcis Lange-Bertalot in Rumrich, Lange-Bertalot & Rumrich, 2000 Navicula vitabunda Hustedt, 1930 Navicula vitiosa Schimanski, 1978 Navicula vulpina Kützing, 1844 Navicula walkeri Sovereign, 1958 Navicula weinzierlii Schimanski, 1975 Navicula wiesneri Lange-Bertalot, 1993 Navicula wildii Lange-Bertalot, 1993 Navicula zanonii Hustedt, 1949
CA, ID, MT, OR, SD, WA, WY CA, ID, MT, OR, WY CA, ID, MT, OR, SD, WA ID, MT MT, ND OR CA, MT, WY CA, ID, MT, OR, WY CA, ID, MT, ND, OR, SD, WA, WY MT
Naviculadicta elorantana Lange-Bertalot in Lange-Bertalot & Metzeltin, 1996 Naviculadicta ventraloconfusa (Lange-Bertalot) Lange-Bertalot in Lange-Bertalot & Moser, 1994
MT MT
Neidiopsis levanderi (Hustedt) Lange-Bertalot & Metzeltin in Lange-Bertalot & Genkal, 1999 Neidiopsis wulfii (Petersen) Lange-Bertalot in Lange-Bertalot & Genkal, 1999
MT MT
Neidium affine (Ehrenberg) Pfitzer, 1871 Neidium affine var. humerus Reimer in Patrick & Reimer, 1966 Neidium affine var. longiceps (Gregory) Cleve, 1894 Neidium affine var. undulatum (Grunow) Cleve, 1894 Neidium alpinum Hustedt, 1943 Neidium amphigomphus (Ehrenberg) Pfitzer, 1871 Neidium ampliatum (Ehrenberg) Krammer in Krammer & Lange-Bertalot, 1985 Neidium apiculatum Reimer, 1959 Neidium binodeforme Krammer in Krammer & Lange-Bertalot, 1985 Neidium bisulcatum (Lagerstedt) Cleve, 1894 Neidium dubium (Ehrenberg) Cleve, 1894 Neidium hercynicum Mayer, 1915 Neidium incurvum var. directum Cleve-Euler, 1955 Neidium iridis (Ehrenberg) Cleve, 1894 Neidium ladogensis (Cleve) Foged, 1952 Neidium productum (W. Smith) Cleve, 1894 Neidium septentrionalis Cleve-Euler, 1939 Neidium temperei Reimer, 1959 Neidium testa Carter & Bailey-Watts, 1981
CA, ID, MT, WY MT MT MT MT MT CA, ID, MT, ND, OR, WA, WY MT CA, ID, MT, OR, SD, WY ID, MT, OR CA, ID, MT, NV, WY CA, ID, MT MT CA, ID, MT MT CA, ID, MT MT MT MT
Nitzschia acicularioides Hustedt, 1959 Nitzschia acicularis (Kützing) W. Smith, 1853 Nitzschia acidoclinata Lange-Bertalot, 1976 Nitzschia acula (Kützing) Hantzsch in Rabenhorst, 1861-1882
CA, MT, WY CA, ID, MT, ND, NV, OR, SD, WA, WY CA, ID, MT, NV, OR, SD, WA, WY MT
26
L.L. BAHLS
Checklist cont. Nitzschia aequorea Hustedt, 1939 Nitzschia agnita Hustedt, 1957 Nitzschia alpina Hustedt, 1943 Nitzschia alpinobacillum Lange-Bertalot, 1993 Nitzschia amphibia Grunow, 1862 Nitzschia amphibia f. frauenfeldii (Grunow) Lange-Bertalot in Lange-Bertalot & Krammer, 1987 Nitzschia amphibioides Hustedt, 1942 Nitzschia amplectens Hustedt, 1957 Nitzschia angustata (W. Smith) Grunow, 1880 Nitzschia angustatula Lange-Bertalot in Lange-Bertalot & Krammer, 1987 Nitzschia angustiforaminata Lange-Bertalot, 1980 Nitzschia archibaldii Lange-Bertalot, 1980 Nitzschia aurariae Cholnoky, 1965 Nitzschia austriaca Hustedt, 1959 Nitzschia bacilliformis Hustedt, 1922 Nitzschia bacillum Hustedt, 1922 Nitzschia behrei Hustedt, 1959 Nitzschia bergii Cleve-Euler, 1952 Nitzschia bremensis Hustedt in Schmidt et al., 1921 Nitzschia brevissima Grunow in Van Heurck, 1881 Nitzschia bryophila (Hustedt) Hustedt, 1943 Nitzschia bulnheimiana (Rabenhorst) H. L. Smith, 1876-1888 Nitzschia capitellata Hustedt in Schmidt et al., 1922 Nitzschia clausii Hantzsch, 1860 Nitzschia communis Rabenhorst, 1860 Nitzschia commutata Grunow in Cleve & Grunow, 1880 Nitzschia commutatoides Lange-Bertalot in Lange-Bertalot & Krammer, 1987 Nitzschia compressa var. balatonis (Grunow) Lange-Bertalot in Lange-Bertalot & Krammer, 1987 Nitzschia compressa var. vexans (Grunow) Lange-Bertalot in Lange-Bertalot & Krammer, 1987 Nitzschia confinis Hustedt, 1949 Nitzschia dealpina Lange-Bertalot & Hofmann in Lange-Bertalot, 1993 Nitzschia desertorum Hustedt, 1949 Nitzschia dippelii Grunow, 1882 Nitzschia dissipata (Kützing) Grunow, 1862 Nitzschia dissipata var. media (Hantzsch) Grunow in Van Heurck, 1881 Nitzschia diversa Hustedt, 1959 Nitzschia draveillensis Coste & Ricard, 1980 Nitzschia dubia W. Smith, 1853 Nitzschia eglei Lange-Bertalot in Lange-Bertalot & Krammer, 1987 Nitzschia elegantula Grunow in Van Heurck, 1881 Nitzschia fasciculata (Grunow) Grunow in Van Heurck, 1881 Nitzschia filiformis (W. Smith) Van Heurck, 1896 Nitzschia flexa Schumann, 1862 Nitzschia flexoides Geitler, 1968 Nitzschia fonticola (Grunow) Grunow in Van Heurck, 1881 Nitzschia fonticoloides Sovereign, 1958 Nitzschia fossilis (Grunow) Grunow in Van Heurck, 1881 Nitzschia frustulum (Kützing) Grunow in Cleve & Grunow, 1880 Nitzschia frustulum var. subsalina Hustedt, 1930 Nitzschia fruticosa Hustedt, 1957
CA, ID, MT, ND, NV, OR, WA, WY CA, ID, MT, ND, NV, OR, SD, WA, WY CA, ID, MT, NV, OR, SD, WA, WY CA, ID, MT, OR, SD, WY CA, ID, MT, ND, NV, OR, SD, WA, WY MT, OR CA, MT, WY MT CA, ID, MT, ND, OR, SD, WA, WY CA, ID, MT, ND, OR, SD, WA, WY ID, MT, ND, OR, SD, WY CA, ID, MT, ND, NV, OR, SD, WA, WY CA, ID, MT, ND, NV, SD, WY MT ID, MT CA, ID, MT, OR, SD, WA, WY MT CA, ID, MT, ND, SD, WY SD CA, ID, WA, WY CA, ID, MT, OR, SD, WY CA, MT, ND CA, ID, MT, ND, NV, OR, SD, WA, WY CA, MT, ND, WY CA, ID, MT, ND, NV, OR, SD, WA, WY CA, MT, ND, OR, SD MT CA CA, MT, ND ID, MT, WA ND, WY CA, ID, MT, ND, SD, WY MT CA, ID, MT, ND, NV, OR, SD, WA, WY MT, WY MT, OR, WA, WY CA, ID, MT, ND, SD, WA, WY CA, MT, WA, WY MT, ND CA, ID, MT, SD, WY MT CA, ID, MT, ND, NV, OR, SD, WY ID, MT, ND ID, MT, SD CA, ID, MT, ND, NV, OR, SD, WA, WY CA, MT CA, ID, MT, ND, OR, SD, WA, WY CA, ID, MT, ND, NV, OR, SD, WA, WY CA, MT, ND, OR, WY MT
A CHECKLIST OF DIATOMS FROM INLAND WATERS OF THE NORTHWESTERN U.S.
27
Checklist cont. Nitzschia gisela Lange-Bertalot in Lange-Bertalot & Krammer, 1987 Nitzschia graciliformis Lange-Bertalot & Simonsen, 1978 Nitzschia gracilis Hantzsch, 1860 Nitzschia hantzschiana Rabenhorst, 1860 Nitzschia heufleriana Grunow, 1862 Nitzschia homburgiensis Lange-Bertalot, 1978 Nitzschia ignorata Krasske, 1929 Nitzschia incognita Legler & Krasske, 1940 Nitzschia inconspicua Grunow, 1862 Nitzschia innominata Sovereign, 1958 Nitzschia intermedia Hantzsch ex Cleve & Grunow, 1880 Nitzschia lacunarum Hustedt in Schmidt et al., 1922 Nitzschia lacuum Lange-Bertalot, 1980 Nitzschia lanceola var. minutula Grunow in Cleve & Grunow, 1880 Nitzschia lanceolata W. Smith, 1853 Nitzschia lancettula Sovereign, 1958 Nitzschia leistikowii Lange-Bertalot, 1980 Nitzschia liebetruthii Rabenhorst, 1864 Nitzschia liebetruthii var. major Cleve & Möller, 1879 Nitzschia linearis (Agardh) W. Smith, 1853 Nitzschia littorea Grunow in Van Heurck, 1881 Nitzschia lorenziana Grunow in Cleve & Grunow, 1880 Nitzschia lorenziana var. subtillis Grunow in Cleve & Grunow, 1880 Nitzschia microcephala Grunow, 1880 Nitzschia modesta Hustedt ex Brendemuhl, 1949 Nitzschia nana Grunow in Van Heurck, 1881 Nitzschia normannii Grunow, 1880 Nitzschia obtusa W. Smith, 1853 Nitzschia ovalis Arnott in Cleve & Grunow, 1880 Nitzschia palea (Kützing) W. Smith, 1856 Nitzschia palea var. tenuirostris Grunow in Van Heurck, 1881 Nitzschia paleacea Grunow in Van Heurck, 1881 Nitzschia paleaeformis Hustedt, 1950 Nitzschia palustris Hustedt, 1934 Nitzschia pellucida Grunow in Cleve & Grunow, 1880 Nitzschia perminuta (Grunow in Van Heurck) Peragallo, 1903 Nitzschia perspicua Cholnoky, 1960 Nitzschia pseudofonticola Hustedt, 1942 Nitzschia pumila Hustedt, 1954 Nitzschia pura Hustedt, 1954 Nitzschia pusilla Grunow, 1862 Nitzschia radicula Hustedt, 1942 Nitzschia rautenbachiae Cholnoky, 1957 Nitzschia recta Hantzsch ex Rabenhorst, 1862 Nitzschia rectiformis Hustedt, 1943 Nitzschia reversa W. Smith, 1853 Nitzschia rhombica Hustedt, 1955 Nitzschia rosenstockii Lange-Bertalot, 1980 Nitzschia scalaris (Ehrenberg) W. Smith, 1853 Nitzschia scalpelliformis Grunow in Cleve & Grunow, 1880
SD CA, ID, MT, ND, OR, SD, WY CA, ID, MT, ND, NV, OR, SD, WA, WY CA, ID, MT, NV, OR, SD, WA, WY CA, ID, MT, ND, NV, OR, SD, WA, WY ID, MT, SD MT CA, ID, MT, ND, NV, OR, SD, WA, WY CA, ID, MT, ND, NV, OR, SD, WA, WY CA, ID, MT, OR, WY CA, ID, MT, ND, NV, OR, SD, WA, WY ID, MT, SD, WY CA, ID, MT, NV, OR, SD, WA, WY CA ID, MT CA, ID, MT, OR CA, MT, OR, WY CA, ID, MT, ND, NV, OR, SD, WY MT CA, ID, MT, ND, NV, OR, SD, WA, WY CA, MT, WY CA, MT, SD, WY MT CA, ID, MT, ND, NV, OR, SD, WY ID, MT, OR, WA, WY CA, MT, ND, WY MT, WA MT, ND, SD, WY CA, MT CA, ID, MT, ND, NV, OR, SD, WA, WY MT CA, ID, MT, ND, NV, OR, SD, WA, WY ID, MT, WA MT MT, WA CA, ID, MT, ND, NV, OR, SD, WA, WY MT, ND, OR, SD, WY ID, WA, WY CA, ID, MT, WA, WY CA, ID, MT, OR, SD, WA, WY CA, ID, MT, ND, NV, OR, SD, WA, WY CA, ID, MT, ND, OR, SD, WY MT CA, ID, MT, ND, NV, OR, SD, WA, WY ID CA, MT, ND, SD, WY OR CA, ID, MT MT CA, MT, WY
28
L.L. BAHLS
Checklist cont. Nitzschia sigma (Kützing) W. Smith, 1853 Nitzschia sigmaformis Hustedt, 1955 Nitzschia sigmoidea (Nitzsch) W. Smith, 1853 Nitzschia siliqua Archibald, 1966 Nitzschia sinuata (Thwaites in W. Smith) Grunow in Cleve & Grunow, 1880 Nitzschia sinuata var. delognei (Grunow in Van Heurck) Lange-Bertalot, 1980 Nitzschia sinuata var. tabellaria (Grunow) Grunow in Van Heurck, 1881 Nitzschia sociabilis Hustedt, 1957 Nitzschia solita Hustedt, 1953 Nitzschia subacicularis Hustedt in Schmidt et al., 1922 Nitzschia subcohaerens var. scotica (Grunow in Van Heurck) Van Heurck, 1896 Nitzschia subinflata Hustedt in Schmidt et al., 1922 Nitzschia sublinearis Hustedt in Schmidt et al., 1921 Nitzschia subtilioides Hustedt, 1959 Nitzschia subtilis (Kützing) Grunow in Cleve & Grunow, 1880 Nitzschia subvitrea Hustedt, 1922 Nitzschia suchlandti Hustedt, 1943 Nitzschia supralitorea Lange-Bertalot, 1979 Nitzschia tarda Hustedt, 1949 Nitzschia terrestris (Petersen) Hustedt, 1934 Nitzschia thermaloides Hustedt, 1955 Nitzschia tropica Hustedt, 1949 Nitzschia tubicola Grunow in Cleve & Grunow, 1880 Nitzschia umbonata (Ehrenberg) Lange-Bertalot, 1978 Nitzschia valdecostata Lange-Bertalot & Simonsen, 1978 Nitzschia valdestriata Aleem & Hustedt, 1951 Nitzschia vermicularis (Kützing) Hantzsch in Rabenhorst, 1848-1860 Nitzschia vitrea Norman, 1861 Nitzschia wuellerstorffii Lange-Bertalot in Lange-Bertalot & Krammer, 1987
CA, ID, MT, ND, NV, OR, SD, WY WY CA, ID, MT, ND, NV, OR, SD, WA, WY CA, ID, MT, SD, WY CA, ID, MT, SD, WA, WY CA, MT CA CA, ID, MT, ND, NV, OR, SD, WA, WY CA, ID, MT, ND, NV, OR, SD, WY CA, ID, MT, ND, WY SD MT CA, ID, MT, OR, SD, WA, WY WY CA, MT, ND, OR, SD, WY MT ID CA, ID, MT, ND, NV, OR, SD, WA, WY MT MT MT, WY MT, OR, WY CA, ID, MT, WA CA, ID, MT, ND, NV, OR, SD, WY CA, ID, MT, ND, NV, OR, SD, WY CA, ID MT, ND, OR, SD, WY CA, ID, MT, ND, NV, OR, SD, WA, WY ID, MT, ND, WY CA, ID, MT, OR, WA, WY
Nupela carolina Potapova & Clason in Potapova et al., 2003 Nupela lapidosa (Krasske) Lange-Bertalot, 1999 Nupela neglecta Ponader, Lowe & Potapova in Potapova et al., 2003 Nupela silvahercynia (Lange-Bertalot) Lange-Bertalot in Lange-Bertalot & Metzeltin, 1996
MT CA, ID, MT, OR, WA CA, MT MT, OR
Opephora ansata Hohn & Hellerman, 1963 Opephora olsenii M. Møller, 1950 Opephora pacifica (Grunow) Petit, 1888
MT CA, ID, MT, OR, SD, WY MT
Orthoseira dendroteres (Ehrenberg) Round, Crawford & Mann, 1990 Orthoseira roeseana (Rabenhorst) O'Meara, 1876
ID, MT, WA ID, MT, WA
Parlibellus crucicula (W. Smith) Witkowski, Lange-Bertalot & Metzeltin, 2000 Parlibellus protracta (Grunow) Witkowski, Lange-Bertalot & Metzeltin, 2000
MT CA, ID, MT, OR, WA, WY
Pinnularia acrosphaeria Rabenhorst, 1853 Pinnularia amabilis Krammer, 2000 Pinnularia appendiculata (Agardh) Cleve, 1895 Pinnularia biceps Gregory, 1856 Pinnularia borealis Ehrenberg, 1843
MT MT CA, ID, MT, OR, WA, WY ID, MT, WA CA, ID, MT, ND, OR SD, WA, WY
A CHECKLIST OF DIATOMS FROM INLAND WATERS OF THE NORTHWESTERN U.S.
29
Checklist cont. Pinnularia borealis var. lanceolata Hustedt in Schmidt et al., 1934 Pinnularia brevicostata Cleve, 1891 Pinnularia decrescens (Grunow) Krammer, 2000 Pinnularia distinguenda (Cleve) Cleve, 1895 Pinnularia divergens W. Smith, 1853 Pinnularia divergens var. media Krammer, 2000 Pinnularia divergentissima (Grunow in Van Heurck) Cleve, 1895 Pinnularia esox Ehrenberg, 1843 Pinnularia gibba Ehrenberg, 1843 Pinnularia gibbiformis Krammer, 1992 Pinnularia hemiptera (Kützing) Rabenhorst, 1853 Pinnularia ignobilis (Krasske) Cleve-Euler, 1955 Pinnularia interrupta W. Smith, 1853 Pinnularia krookii (Grunow) Cleve, 1891 Pinnularia kuetzingii Krammer, 1992 Pinnularia lagerstedtii (Cleve) Cleve-Euler, 1934 Pinnularia lata (Brébisson) W. Smith, 1853 Pinnularia legumen Ehrenberg, 1843 Pinnularia lundii Hustedt, 1954 Pinnularia lundii var. linearis Krammer, 2000 Pinnularia macilenta Ehrenberg, 1843 Pinnularia maior (Kützing) Rabenhorst, 1833 Pinnularia marchica Schönfelder, 2000 Pinnularia mesogongyla var. interrupta Cleve, 1891 Pinnularia mesolepta (Ehrenberg) W. Smith, 1853 Pinnularia microstauron (Ehrenberg) Cleve, 1891 Pinnularia microstauron var. nonfasciata Krammer, 2000 Pinnularia microstauron var. rostrata Krammer, 2000 Pinnularia neomajor Krammer, 1992 Pinnularia nobilis (Ehrenberg) Ehrenberg, 1843 Pinnularia nodosa (Ehrenberg) W. Smith, 1856 Pinnularia obscura Krasske, 1932 Pinnularia peracuminata Krammer, 2000 Pinnularia pisciculus Ehrenberg, 1843 Pinnularia polyonca (Brébisson in Kützing) W. Smith, 1856 Pinnularia pseudogibba Krammer, 1992 Pinnularia pulchra Østrup, 1897 Pinnularia rhombarea Krammer in Metzeltin & Lange-Bertalot, 1998 Pinnularia rupestris Hantzsch in Rabenhorst, 1861 Pinnularia schoenfelderi Krammer, 1992 Pinnularia schroederii (Hustedt) Krammer in Krammer & Lange-Bertalot, 1985 Pinnularia schwabei Krasske, 1939 Pinnularia scotica Krammer, 2000 Pinnularia septentrionalis Krammer, 2000 Pinnularia silvatica Petersen, 1935 Pinnularia stomatophora (Grunow in Schmidt et al.) Cleve, 1895 Pinnularia streptoraphe Cleve, 1891 Pinnularia subanglica Krammer, 2000 Pinnularia subcapitata Gregory, 1856 Pinnularia subcommutata Krammer, 1992
OR MT MT ID ID, MT, WA ID CA, ID, MT, OR, WY MT CA, ID, MT, OR, SD, WA, WY ID, MT CA, MT ID, MT, ND, OR, SD, WY CA, ID, MT, WY MT ID, MT, WA MT ID, MT ID, MT, WA CA, MT MT MT ID, MT, WA WA MT ID, MT, WA CA, ID, MT, ND, OR SD, WA, WY MT MT MT CA ID, MT, OR, WA, WY CA, ID, MT, OR, WA, WY ID MT MT ID, MT MT, WA MT WA WA CA ID, MT ID ID, MT ID, WA ID, MT, OR, WY ID, MT, ND MT CA, ID, MT, ND, OR, SD, WA, WY WA
30
L.L. BAHLS
Checklist cont. Pinnularia subgibba Krammer, 1992 Pinnularia subgigas Krammer, 2000 Pinnularia subrostrata (A. Cleve) Cleve-Euler, 1955 Pinnularia subrupestris Krammer, 1992 Pinnularia sudetica (Hilse in Rabenhorst) M. Peragallo, 1903 Pinnularia transversa (Schmidt in Schmidt et al.) Mayer, 1939 Pinnularia turbulenta (Cleve-Euler) Krammer, 2000 Pinnularia viridiformis Krammer, 1992 Pinnularia viridis (Nitzsch) Ehrenberg, 1843
ID, MT, WA MT MT ID, MT ID, MT, OR WA MT MT, WA CA, ID, MT, ND, OR, SD, WA, WY
Placoneis abiskoensis (Hustedt) Lange-Bertalot & Metzeltin in Metzeltin & Witkowski, 1996 Placoneis clementioides (Hustedt) Cox, 1987 Placoneis clementis (Grunow) Cox, 1987 Placoneis elginensis (Gregory) Cox, 1987 Placoneis exigua (Gregory) Mereschkowsky, 1903 Placoneis explanata (Hustedt) Mayama in Mayama & Kawashima, 1998 Placoneis gastrum (Ehrenberg) Mereschkowsky, 1903 Placoneis placentula (Ehrenberg) Mereschkowsky, 1903 Placoneis pseudanglica Cox, 1987
MT, ND CA, ID, MT, OR, WY CA, ID, MT, NV, OR, WA, WY CA, ID, MT, NV, OR, SD, WA, WY CA, ID, MT, WY MT, WA, WY ID, MT, WY CA, ID, MT, ND, OR, WY CA, ID, MT, ND, NV, SD, WY
Plagiotropis arizonica Czarnecki & Blinn, 1978 Plagiotropis lepidoptera (Gregory) Kuntze, 1898 Plagiotropis lepidoptera var. proboscidea (Cleve) Reimer in Patrick & Reimer, 1975
MT, SD MT, ND, SD, WY CA, MT
Planothidium apiculatum (Patrick) Lange-Bertalot, 1999 Planothidium biporomum (Hohn & Hellerman) Lange-Bertalot, 1999 Planothidium calcar (Cleve) Round & Bukhtiyarova, 1996 Planothidium daui (Foged) Lange-Bertalot, 1999 Planothidium delicatulum (Kützing) Round & Bukhtiyarova, 1996 Planothidium dubium (Grunow) Round & Bukhtiyarova, 1996 Planothidium ellipticum (Cleve) Round & Bukhtiyarova, 1996 Planothidium engelbrechtii (Cholnoky) Round & Bukhtiyarova, 1996 Planothidium frequentissimum (Lange-Bertalot) Round & Bukhtiyarova, 1996 Planothidium granum (Hohn & Hellerman) Lange-Bertalot, 1999 Planothidium haukianum (Grunow) Round & Bukhtiyarova, 1996 Planothidium haynaldii (Schaarschmidt) Lange-Bertalot, 1999 Planothidium joursacense (Héribaud) Lange-Bertalot, 1999 Planothidium lanceolatoide (Sovereign) Lange-Bertalot, 1999 Planothidium lanceolatum (Brébisson ex Kützing) Lange-Bertalot, 1999 Planothidium minutissimum (Krasske) Lange-Bertalot, 1999 Planothidium oestrupii (Cleve-Euler) Round & Bukhtiyarova, 1996 Planothidium peragalli (Brun & Héribaud) Round & Bukhtiyarova, 1996 Planothidium robustius Lange-Bertalot, 1999 Planothidium robustum (Hustedt) Lange-Bertalot, 1999 Planothidium rostratum (Østrup) Lange-Bertalot, 1999 Planothidium stewartii (Patrick) Lange-Bertalot, 2004
CA, ID, MT CA, ID, MT, OR, WA, WY CA, ID, MT, OR, WA CA, ID, MT, WY CA, ID, MT, ND, OR, SD, WY CA, ID, MT, OR, SD, WA, WY SD CA, ID, MT, OR, WY CA, ID, MT, ND, OR, SD, WA, WY CA, ID, MT, OR, WY CA, MT, ND, SD, WA, WY CA, ID, MT, SD, WY MT MT, OR, WY CA, ID, MT, OR, SD, WA, WY CA, MT, OR ID, MT, OR, WY CA, ID, MT, OR, WY ID WY CA, ID, MT, SD, WY ID, OR, WY
Platessa conspicua (Mayer) Lange-Bertalot, 2004 Platessa hustedtii (Krasske) Lange-Bertalot, 2004 Platessa montana (Krasske) Lange-Bertalot, 2004
CA, ID, MT, OR, WA, WY CA, WA MT
A CHECKLIST OF DIATOMS FROM INLAND WATERS OF THE NORTHWESTERN U.S.
31
Checklist cont. Pleurosigma angulatum (Quekett) W. Smith, 1852 Pleurosigma delicatulum W. Smith, 1852 Pleurosigma elongatum W. Smith, 1852 Pleurosigma salinarum (Grunow) Grunow in Cleve & Grunow, 1880
CA CA, MT, ND, NV, OR, SD, WY MT, SD, WY CA
Pleurosira laevis (Ehrenberg) Compère, 1982
CA, ID, MT, OR, WY
Proshkinia bulnheimii (Grunow) Karayeva, 1978 Proshkinia complanata (Grunow) Mann, 1990
MT SD
Psammothidium abundans f. rosenstockii (Lange-Bertalot in Lange-Bertalot & Krammer) Bukhtiyarova in Bukhtiyarova & Round, 1996
MT
Psammothidium altaicum (Poretzky) Bukhtiyarova in Bukhtiyarova & Round, 1996 Psammothidium bioretii (Germain) Bukhtiyarova & Round, 1996 Psammothidium chlidanos (Hohn & Hellerman) Lange-Bertalot, 1999 Psammothidium daonense (Lange-Bertalot in Lange-Bertalot & Krammer) Lange-Bertalot, 1999 Psammothidium didymum (Hustedt) Bukhtiyarova & Round, 1996 Psammothidium grischunum (Wuthrich) Bukhtiyarova & Round, 1996 Psammothidium helveticum (Hustedt) Bukhtiyarova & Round, 1996 Psammothidium lacus-vulcani (Lange-Bertalot & Krammer) Bukhtiyarova in Bukhtiyarova & Round, 1996
MT CA, ID, MT, NV, OR, WA, WY CA, ID, MT, OR, WA, WY CA, ID, MT, OR, WA, WY ID, MT, WA CA, ID, MT, OR, WA, WY CA, MT, OR, WY MT, OR
Psammothidium lauenbergianum (Hustedt) Bukhtiyarova & Round, 1996 Psammothidium levanderi (Hustedt) Bukhtiyarova & Round, 1996 Psammothidium marginulatum (Grunow) Bukhtiyarova & Round, 1996 Psammothidium rossii (Hustedt) Bukhtiyarova & Round, 1996 Psammothidium sacculum (Carter) Bukhtiyarova in Bukhtiyarova & Round, 1996 Psammothidium scoticum (Flower & Jones) Bukhtiyarova & Round, 1996 Psammothidium subatomoides (Hustedt) Bukhtiyarova & Round, 1996 Psammothidium ventralis (Krasske) Bukhtiyarova & Round, 1996
ID, MT, OR, WY MT, OR, WA CA, ID, MT, OR, WA, WY CA, ID, MT, OR, WY ID, MT, WA MT CA, ID, MT, OR, WA, WY CA, ID, MT, WA, WY
Pseudostaurosira brevistriata (Grunow in Van Heurck) Williams & Round, 1987 Pseudostaurosira brevistriata var. inflata (Pantocsek) Edlund, 1994 Pseudostaurosira elliptica (Schumann) Edlund, Morales & Spaulding, 2006 Pseudostaurosira parasitica (W. Smith) Morales, 2003 Pseudostaurosira parasitica var. subconstricta (Grunow) Morales, 2003 Pseudostaurosira pseudoconstruens (Marciniak) Williams & Round, 1987 Pseudostaurosira robusta (Fusey) Williams & Round, 1987
CA, ID, MT, ND, NV, OR, SD, WA, WY CA, MT CA, MT, ND, SD, WY CA, ID, MT, ND, NV, OR, SD, WA, WY CA, ID, MT ID, MT, OR, WY CA, ID, MT
Puncticulata bodanica (Grunow in Schneider) Håkansson, 2002 Puncticulata comta (Ehrenberg) Håkansson, 2002 Puncticulata glabriuscula (Grunow) Håkansson, 2002 Puncticulata praetermissa (Lund) Håkansson, 2002 Puncticulata radiosa (Lemmermann) Håkansson, 2002
CA, ID, MT, ND, OR, WY MT MT, WY MT MT
Reimeria sinuata (Gregory) Kociolek & Stoermer, 1987 Reimeria sinuata f. antiqua (Grunow) Kociolek & Stoermer, 1987 Reimeria uniseriata Sala, Guerrero & Ferrario, 1993
CA, ID, MT, ND, NV, OR, SD, WA, WY MT CA, MT, ND, OR, SD, WY
Rhoicosphenia abbreviata (C. Agardh) Lange-Bertalot, 1980
CA, ID, MT, ND, OR, SD, WA, WY
32
L.L. BAHLS
Checklist cont. Rhopalodia acuminata Krammer in Lange-Bertalot & Krammer, 1987 Rhopalodia brebissonii Krammer in Lange-Bertalot & Krammer, 1987 Rhopalodia constricta (W. Smith) Krammer in Lange-Bertalot & Krammer, 1987 Rhopalodia gibba (Ehrenberg) O. Müller, 1895 Rhopalodia gibberula (Ehrenberg) O. Müller, 1895 Rhopalodia musculus (Kützing) O. Müller, 1900 Rhopalodia operculata (C. Agardh) Håkansson, 1979 Rhopalodia rupestris (W. Smith) Krammer in Krammer & Lange-Bertalot, 1987
CA, ID, MT, OR, WY CA, MT, ND, NV, OR, SD, WY ID, MT, SD, WY CA, ID, MT, ND, NV, OR, SD, WA, WY CA, ID, MT, ND, SD, WY CA, ID, MT, WY CA, ID, MT, ND, OR, SD, WY MT
Rossithidium linearis (W. Smith) Round & Bukhtiyarova, 1996 Rossithidium petersenii (Hustedt) Round & Bukhtiyarova, 1996 Rossithidium pusillum (Grunow) Round & Bukhtiyarova, 1996
CA, ID, MT, OR, WA ID, MT, OR CA, ID, MT, OR, SD, WA, WY
Scoliopleura peisonis Grunow, 1860
MT
Sellaphora alastos (Hohn & Hellerman) Lange-Bertalot & Metzeltin, 1996 Sellaphora americana (Ehrenberg) Mann, 1989 Sellaphora bacillum (Ehrenberg) Mann, 1989 Sellaphora disjuncta (Hustedt) Mann, 1989 Sellaphora hustedtii (Krasske) Lange-Bertalot & Werum in Werum & Lange-Bertalot, 2004 Sellaphora laevissima (Kützing) Mann, 1989 Sellaphora mutata (Krasske) Lange-Bertalot in Lange-Bertalot et al., 1996 Sellaphora parapupula Lange-Bertalot in Lange-Bertalot & Metzeltin, 1996 Sellaphora pupula (Kützing) Mereschkowsky, 1902 Sellaphora pupula var. densestriata (Kützing) Lange-Bertalot & Metzeltin, 1996 Sellaphora pupula var. elliptica (Hustedt) Bukhtiyarova, 1995 Sellaphora pupula f. rostrata (Hustedt) Bukhtiyarova, 1995 Sellaphora rectangularis (Gregory) Lange-Bertalot & Metzeltin, 1996 Sellaphora seminulum (Grunow) Mann 1989
MT ID, MT CA, ID, MT, OR, WY ID, MT CA, ID, MT, OR, SD, WA, WY CA, ID, MT, ND, NV, OR, SD, WA, WY ID, MT, WY MT CA, ID, MT, ND, NV, OR, SD, WA, WY MT MT MT MT CA, ID, MT, ND, OR, SD, WA, WY
Simonsenia delognei (Grunow) Lange-Bertalot, 1979
CA, ID, MT, ND, NV, OR, SD, WA, WY
Skeletonema potamos (Weber) Hasle, 1976 Skeletonema subsalsum (Cleve-Euler) Bethge, 1928
MT, WA OR
Stauroforma exiguiformis (Lange-Bertalot) Flower, Jones & Round, 1996
CA, ID, MT, OR, WA, WY
Stauroneis acuta W. Smith, 1853 Stauroneis agrestis Petersen, 1915 Stauroneis anceps Ehrenberg, 1843 Stauroneis borrichii (Petersen) Lund, 1946 Stauroneis circumborealis Lange-Bertalot & Krammer in Lange-Bertalot & Genkal, 1999 Stauroneis fluminea Patrick & Freese, 1961 Stauroneis gracilior Reichardt, 1995 Stauroneis gracilis Ehrenberg, 1843 Stauroneis gracillima Hustedt, 1943 Stauroneis heinii Lange-Bertalot & Krammer in Lange-Bertalot & Genkal, 1999 Stauroneis kriegeri Patrick, 1945 Stauroneis lapidicola Petersen, 1928 Stauroneis lauenburgiana Hustedt, 1950
MT CA, ID, MT CA, ID, MT, ND, OR, WA, WY MT, OR MT MT MT ID, MT MT MT CA, ID, MT, OR, WA, WY MT MT
A CHECKLIST OF DIATOMS FROM INLAND WATERS OF THE NORTHWESTERN U.S.
33
Checklist cont. Stauroneis legumen (Ehrenberg) Kützing, 1844 Stauroneis lineolata Ehrenberg, 1843 Stauroneis montana Krasske ex Hustedt, 1930 Stauroneis neohyalina Lange-Bertalot & Krammer in Lange-Bertalot & Metzeltin, 1996 Stauroneis nobilis var. baconiana (Stodder) Reimer, 1961 Stauroneis obtusa Lagerstedt, 1873 Stauroneis phoenicenteron (Nitzsch) Ehrenberg, 1843 Stauroneis prominula (Grunow in Cleve & Möller; Grunow ex Cleve) Hustedt, 1959 Stauroneis pseudosubobtusoides Germain, 1980 Stauroneis siberica (Grunow in Cleve & Grunow) Lange-Bertalot & Krammer in Lange-Bertalot & Metzeltin, 1996 Stauroneis smithii Grunow, 1860 Stauroneis smithii var. incisa Pantocsek, 1902 Stauroneis tackei (Hustedt) Krammer & Lange-Bertalot, 1985 Stauroneis thermicola (Petersen) Lund, 1946 Stauroneis undata Hustedt, 1939 Stauroneis wislouchii Poretzky & Anissimova, 1933
MT MT ID MT MT CA, ID, MT CA, ID, MT, ND, OR, SD, WA, WY ID, MT OR CA, ID, MT, ND, NV, OR, WA, WY MT CA, ID, MT, ND, NV, OR, SD, WY CA, ID, MT, WA, WY ID MT
Staurosira construens Ehrenberg, 1843 Staurosira construens var. binodis (Ehrenberg) Hamilton in Hamilton et al., 1992 Staurosira construens var. pumila (Grunow) Kingston, 2000 Staurosira venter (Ehrenberg) Cleve & Möller, 1879
CA, ID, MT, ND, NV, OR, SD, WA, WY CA, ID, MT, OR CA CA, ID, MT, ND, WY
Staurosirella ansata (Hohn & Hellerman) Kingston, 2000 Staurosirella dubia (Grunow) Morales & Manoylov, 2006 Staurosirella lapponica (Grunow in Van Heurck) Williams & Round, 1987 Staurosirella leptostauron (Ehrenberg) Williams & Round, 1987 Staurosirella martyi (Héribaud) Morales & Manoylov, 2006 Staurosirella oldenburgiana (Hustedt) Morales, 2005 Staurosirella ovata Morales in Morales & Manoylov, 2006 Staurosirella pinnata (Ehrenberg) Williams & Round, 1987 Staurosirella rhomboides (Grunow) Morales & Manoylov, 2006 Staurosirella subcapitata (Frenguelli) Morales in Morales & Manoylov, 2006 Staurosirella subrobusta Morales in Morales & Manoylov, 2006
CA CA, MT CA, ID, MT, NV CA, ID, MT, NV, OR, SD, WA, WY CA, ID, MT, OR, WA, WY MT, OR CA, MT CA, ID, MT, ND, OR, SD, WA, WY CA, MT MT MT
Stenopterobia anceps (Lewis) Brébisson ex Van Heurck, 1896 Stenopterobia curvula (W. Smith) Krammer in Lange-Bertalot & Krammer, 1987 Stenopterobia delicatissima (Lewis) Van Heurck, 1896
MT ID, MT MT
Stephanocostis chantaicus Genkal & Kuzmina, 1985
MT
Stephanodiscus alpinus Hustedt, 1942 Stephanodiscus hantzschii Grunow in Cleve & Grunow, 1880 Stephanodiscus medius Håkansson, 1986 Stephanodiscus minutulus (Kützing) Cleve & Möller, 1882 Stephanodiscus neoastraea Håkansson & Hickel, 1986 Stephanodiscus niagarae Ehrenberg, 1845 Stephanodiscus oregonica (Ehrenberg) Håkansson, 1986 Stephanodiscus parvus Stoermer & Håkansson, 1984
MT CA, ID, MT, ND, OR, SD, WA, WY CA, ID, MT, ND, OR, SD, WY CA, ID, MT, ND, OR, SD, WA ID, MT CA, ID, MT, ND, OR, SD, WY MT MT, ND, WY
MT
34
L.L. BAHLS
Checklist cont. Surirella angusta Kützing, 1844 Surirella atomus Hustedt, 1955 Surirella bifrons Ehrenberg, 1843 Surirella biseriata Brébisson, 1836 Surirella brebissonii Krammer & Lange-Bertalot, 1987 Surirella brightwellii W. Smith, 1853 Surirella elegans Ehrenberg, 1843 Surirella gracilis (W. Smith) Grunow, 1862 Surirella hoefleri Hustedt, 1959 Surirella iowensis Lowe, 1973 Surirella linearis W. Smith, 1853 Surirella linearis var. constricta Grunow, 1862 Surirella linearis var. helvetica (Brun) Meister, 1912 Surirella minuta Brébisson, 1838 Surirella ovalis Brébisson, 1838 Surirella patella Kützing, 1844 Surirella robusta Ehrenberg, 1840 Surirella spiralis Kützing, 1844 Surirella splendida (Ehrenberg) Kützing, 1844 Surirella striatula Turpin, 1816-1829 Surirella subsalsa W. Smith, 1853 Surirella suecia Grunow in Van Heurck, 1881 Surirella tenera Gregory, 1856
CA, ID, MT, ND, NV, OR, SD, WA, WY MT, SD, WY MT ID, MT CA, ID, MT, ND, NV, OR, SD, WA, WY MT CA, MT CA, ID, MT, ND, WY MT MT CA, ID, MT, ND, OR, SD, WA MT ID, MT, OR, WY CA, ID, MT, ND, NV, OR, SD, WA, WY CA, ID, MT, ND, NV, OR, SD, WY MT CA, MT, OR ID, MT, OR, SD, WA, WY ID, MT, WY MT, SD MT MT CA, ID, MT, ND, OR, SD, WY
Synedra acus Kützing, 1844 Synedra biceps Kützing, 1844 Synedra capitata Ehrenberg, 1836 Synedra cyclopum Brutschy, 1922 Synedra delicatissima W. Smith, 1853 Synedra delicatissima var. angustissima Grunow in Van Heurck, 1881 Synedra famelica Kützing, 1844 Synedra filiformis Grunow in Cleve & Grunow, 1880 Synedra goulardi Brébisson ex Cleve & Grunow, 1880 Synedra mazamaensis Sovereign, 1958 Synedra minuscula Grunow in Van Heurck, 1881 Synedra radians Kützing, 1844 Synedra rumpens Kützing, 1844 Synedra ulna (Nitzsch) Ehrenberg, 1832 Synedra ulna var. chaseana Thomas in Walker & Chase, 1886 Synedra ulna var. contracta Østrup, 1901 Synedra ulna var. danica (Kützing) Grunow in Van Heurck, 1885 Synedra ulna var. oxyrhynchus (Kützing) Van Heurck, 1885 Synedra ulna var. ramesi (Héribaud) Hustedt, 1930 Synedra ulna var. spathulifera (Grunow in Van Heurck) Grunow in Van Heurck, 1885
CA, ID, MT, ND, OR, SD, WA, WY MT MT, SD MT CA, ID, MT, ND, NV, OR, WA, WY MT, WY CA, MT, ND, NV, WY ID, MT MT CA, ID, MT, OR, WA, WY MT MT CA, ID, MT, ND, NV, OR, SD, WA, WY CA, ID, MT, ND, NV, OR, SD, WA, WY MT CA, ID, MT, WY MT MT, WY MT MT
Tabellaria fenestrata (Lyngbye) Kützing, 1844 Tabellaria flocculosa (Roth) Kützing, 1844 Tabellaria quadriseptata Knudson, 1952
ID, MT, WA, WY CA, ID, MT, OR, WA, WY ID, MT
Tabularia fasciculata (Agardh) Williams & Round, 1986
CA, ID, MT, ND, NV, OR, SD, WY
A CHECKLIST OF DIATOMS FROM INLAND WATERS OF THE NORTHWESTERN U.S.
35
Checklist cont.
Tabularia tabulata (Agardh) Snoeijs, 1992 Terpsinoe musica Ehrenberg, 1843
MT CA
Tetracyclus glans (Ehrenberg) Mills, 1935
ID, MT, WA
Thalassiosira baltica (Grunow in Cleve & Grunow) Ostenfeld, 1901 Thalassiosira pseudonana Hasle & Heimdal, 1970 Thalassiosira visurgis Hustedt, 1957 Thalassiosira weissflogii (Grunow) Fryxell & Hasle, 1977
CA, ND CA, ID, MT, ND, OR, SD, WY CA, MT, SD, WY CA, ID, MT, ND, NV, SD, WY
Tryblionella acuminata W. Smith, 1853 Tryblionella acuta (Cleve) Mann in Round, Crawford & Mann, 1990 Tryblionella apiculata Gregory, 1857 Tryblionella calida (Grunow in Cleve & Grunow) Mann in Round, Crawford & Mann, 1990 Tryblionella coarctata (Grunow in Cleve & Grunow) Mann in Round, Crawford & Mann, 1990 Tryblionella compressa (Bailey) Poulin in Poulin et al., 1990 Tryblionella debilis Arnott ex O’Meara, 1873 Tryblionella gracilis W. Smith, 1853 Tryblionella hungarica (Grunow) Mann in Round, Crawford & Mann, 1990 Tryblionella levidensis W. Smith, 1856 Tryblionella littoralis (Grunow in Cleve & Grunow) Mann in Round, Crawford & Mann, 1990 Tryblionella victoriae Grunow, 1862
MT, SD MT CA, ID, MT, ND, NV, OR, SD, WY CA, ID, MT, ND, OR, SD, WY CA CA, ID, MT, ND, SD, WY CA, MT, ND, OR, SD, WA, WY CA, MT, ND, NV, SD, WA, WY CA, ID, MT, ND, NV, OR, SD, WA, WY CA, ID, MT, ND, OR, SD, WA, WY CA, MT, ND, OR, SD, WA, WY CA, ID, MT
Urosolenia eriensis (H. L. Smith) Round & Crawford in Round, Crawford & Mann, 1990
MT