Catalogue of the Heteroptera of
the Palaearctic Region edited by
Berend Aukema & Christian Rieger
Volume 1
Enicocephalomorpha, Dipsocoromorpha, Nepomorpha, Gerromorpha and Leptopodomorpha
CIP-DATA KONINKLUKE BIBLIOTHEEK, DEN HAAG Aukema, B. & Chr. Rieger (editors) Catalogue of Heteroptera of the Palaearctic Region
Catalogue. - With index, references ISBN 90-71912-12-4 Subject headings: Heteroptera I Palaearctic Region I checklist I type material/ distributio references
Published 15 April 1995 by The Netherlands Entomological Society c/o Plantage Middenlaan 64 NL-1018 DH Amsterdam The Netherlands
Printed by Ponsen & Looijen, Wageningen, The Netherlands
Cover: Leptopus hispanus Rambur (Leptopodidae), del. J. Pencart, Montereau. France ii
CONTENTS Preface . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . v Arrangement of the Catalogue . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . ix Introduction . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . ix Depositories (abbreviations) . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . xii Distribution (abbreviations) . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . xv Literature . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . xvi Acknowledgements . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . xxv Enicocephalomorpha . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 Family Aenictopecheidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 Family Enicocephalidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 2 Dipsocoromorpha . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 6 Family Ceratocombidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 6 Family Dipsocoridae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 8 Family Schizopteridae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 10 Nepomorpha . . . . . . . . . Family Nepidae . . . . . Family Belostomatidae . Family Gelastocoridae . Family Ochteridae . . . . Family Corixidae . . . . Family Naucoridae . . . Family Aphelocheiridae Family Notonectidae . . Family Pleidae . . . . . . Family Helotrephidae . .
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13 14 19 23 26 27 57 60 63 73 75
Gerromorpha . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Family Mesoveliidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Family Hebridae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Family Hydrometridae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Family Herrnatobatidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Family Veliidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Family Gerridae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
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77 77 79 83 85 85 96
Leptopodomorpha . . . . Family Aepophilidae Family Saldidae . . . Family Leptopodidae Family Omaniidae
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115 115 116 137 140
References . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 143 Index . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 201
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PREFACE The history of cataloguing Palaearctic Heteroptera started with Herrich-Schaeffer's list of European species (1835a) and his later list of world species (1853). Nearly simultaneously a world list of Heteroptera and Auchenorrhyncha (Dohm, 1859) and a list of European Heteroptera (Baerensprung, 1860) were published. They were followed by Puton's checklists (1869a, 1875, 1886a, and 1899), which were started as lists of European Heteroptera, but the last two editions covered Heteroptera, Auchenorrhyncha and Psylloidea of the entire Palaearctic Region. At the end of the 19th century the publication of a world catalogue of Heteroptera was started by Lethierry & Severin (1893-1896), but it was never completed. The first comprehensive catalogue of Palaearctic Heteroptera was written by V.F. Oshanin and published by the Zoological Museum of St. Petersburg in the years 1906-1910. In fact it was a German translation of an earlier Russian version already fmished in 1892. A shortened version of the catalogue was published in Berlin in 1912. Stichel (1955-1962) subsequently included a checklist of Palaearctic species, with references to original descriptions and synonymies published after Oshanin's catalogue, in his identification keys of European Heteroptera. In 1988 during an international symposium on Entomofaunistics of Central Europe in Kiev T. Vasarhelyi launched the idea of compiling a new catalogue of Palaearctic Heteroptera, to be published in Budapest, and discussed it with I.M. Kerzhner, J. Pericart and P.V. Putshkov. Later that same year a meeting of I.M. Kerzhner, P. Stys and T. Vasarhelyi was organized in Budapest to develop further plans, and it was decided that A. Jansson and I.M. Kerzhner would prepare preliminary texts on Corixidae and Nabidae respectively. Unfortunately T. Vasarhelyi had to withdraw as editor and later on also publication in Hungary became impossible. Not earlier than during the 4th European Congress of Entomology in 1990 in GOdollo, Hungary, the matter was picked up again and discussed during the "Heteuroptera" workshop, where B. Aukema offered to check the possibilities of publication in The Netherlands. After getting the support of the Nederlandse Entomologische Vereniging (N.E.V.) the work was started with B. Aukema and Chr. Rieger as editors. We dedicate this work to the memory of Vasiliy Fedorovich Oshanin (1844-1917), author of the first comprehensive catalogue of Palearctic Heteroptera. Berend Aukema Christian Rieger
PREFACE TO THE FIRST VOLUME With great pleasure we herewith present the first volume of what can be considered the second "Catalogue of Palaearctic Heteroptera•. It contains Enicocephalomorpha and Dipsocoromorpha by I.M. Kerzhner, Nepomorpha by J. Polhemus, A. Jansson and E.V. Kanyukova, Gerromorpha by N.M. Anderson, and Leptopodomorpha by P. Lindskog. In total 25 families of these infraorders are represented in the Palaearctic Region by altogether 647 species and 42 subspecies of 121 genera (table 1).
Numbers of Palaearctic Heteroptera Genera Species Subspecies Enicocephalomorpha Dipsocoromorpha Nepomorpha
Gerromorpha
Leptopodomorpha
Aenictopecheidae Enicocephalidae Ceratocombidae Dipsocoridae Schizopteridae Nepidae Be1ostomatidae Gelastocoridae Ochteridae Corixidae Naucoridae Aphelocheiridae Notonectidae Pleidae Helotrephidae Mesoveliidae Hebridae Hydrometridae Herrnatobatidae Veliidae Gerridae Aepophilidae Saldidae Leptopodidae Omaniidae
Totals
1 6 1 1 6 5 5 1 1 15 7 1 4 2 5 2 4 1 1 20 1 14 4 2
1 15 11 14 9 21 14 4 3 143 9 18 50 6 6 7 24 14 2 64 99 1 99 11 2
121
647
11
12 3 5 1
3 9 5 4
42
Table 1. Numbers of Palaearctic Heteroptera listed in volume 1. The following new synonymies are included. Gerridae: Gerris costae poissoni Wagner & Zimmermann = G. costae arvemenis Poisson. Saldidae: Calacanthia trybomi (J. Sahlberg) = ?Pedeticus variegaJUS Curtis; Chanoscina cocksii (Curtis) = C. geminata f. putoni
Stichel; Micracanthia ornatula (Reuter)
= Sa/du/a
minor Hamid & Sultana; Saldula
arenicola arenicola (Scholtz) = Salda ocellata A. Costa; Saldula fudcola (J. Sahlberg) = Salda riparia Zetterstedt; Saldula opacula (Zetterstedt) = Salda quadrilineata Jakovlev;
vii
Saldula recticollis (Horvath) = S. fuldena Drake & Maa = S. (S.) arsenjevi Vinokurov. In Saldidae the following new combinations are included: Micracanthia bonina (Drake) = Saldula bonina Drake; Micracanthia hasegawai (Cobben) = Saldula hasegawai Cobben; Micracanthia omatula (Reuter) = Salda omatula Reuter.
Special thanks are due to I.M. Kerzhner, St. Petersburg, Russia, and J.T. Polhemus, Englewood, Colorado, USA, who were of great help to the editors in reviewing all contributions to the first volume and for supplying a lot of additional information. L. Y. Zheng and eo-workers, Tianjin, P.R. China, compiled detailed information about the Chinese Heteroptera. The editors keep a computerized database of the Catalogue of Palaearctic Heteroptera. Comments and additional information are welcome at the editorial address: Catalogue of Palaearctic Heteroptera attn B. Aukema & Chr. Rieger Plant Protection Service P.O. Box 9102 NL-6700 HC Wageningen The Netherlands Email:
[email protected]
viii
ARRANGEMENT OF THE CATALOGUE Introduction Heteroptera are used here as a suborder of the Hemiptera, although there are different opinions on the subject (see the discussion in H~nry & Froeschner, 1988; see also Dolling, 1991a). Families are arranged according to Stys & Kerzhner (1975) with some later modifications (Andersen, 1982a: Gerromorpha; Stys & Jansson, 1988: Nepomorpha) as given on the next page. The Catalogue of Palaearctic Heteroptera will be published in five separate volumes as follows. Volume 1: Enicocephalomorpha, Dipsocoromorpha, Nepomorpha, Gerromorpha, and Leptopodomorpha Volume 2: Cimicomorpha (Miridae excluded) Volume 3: Cimicomorpha (Miridae) Volume 4: Pentatomomorpha (Aradoidea, Piesmatoidea and Coreoidea part.) Volume 5: Pentatomomorpha (Stenocephalidae, Coreidae, Alydidae, Rhopalidae, Pentatomoidea and species incertae sedis) Each of the infraorders and families starts with an introductory text including general characteristics of the group and its habits, followed by a listing of the most important references. Suprageneric taxa as a rule are placed in systematic order. Genera, subgenera, species and subspecies are treated alphabetically. In the case of subspecies, however, the nominotypical subspecies is listed first. Family-group names Valid names of superfamilies, families, subfamilies, tribes and subtribes are given with rank, author and year of publication as bold centred headings. Under the heading the valid name and its synonyms are given in chronological order with a reference and the type genus. For extralimital type genera the distribution is briefly indicated. References and additional information (as notes) may be included. Genus-group mimes Valid names of genera and subgenera are given with author and year of publication as bold centred headings. Under the heading the valid name and its synonyms are given in chronological order with a reference, information on original- rank, availability, homonymy, and synonymization or changes of rank. The type species (in original combination) and the method of its fixation are always given. For extralimital type species the distribution is briefly indicated. In the case of replacement names and emendations the type species are not given. Species-group names
Valid names of species and subspecies are given as bold left margined headings. In the case of subspecies the nominotypical subspecies is given first. Extralimital nominotypical subspecies are given between brackets. ix
Classification of Palaearctic families of Heteroptera Infraorder Enicocephalomorpha Superfamily Enicocephaloidea: Aenictopecheidae, Enicocephalidae lnfraorder Dipsocoromorpha Superfamily Dipsocoroidea: Ceratocombidae, Dipsocoridae, Schizopteridae lnfraorder Nepomorpha Superfamily Nepoidea: Nepidae, Belostomatidae Superfamily Corixoidea: Corixidae Superfamily Ochteroidea: Ochteridae, Gelastocoridae Superfamily Naucoroidea: Naucoridae, Aphelocheiridae Superfamily Notonectoidea: Notonectidae Superfamily Pleoidea: Pleidae, Helotrephidae lnfraorder Gerromorpha Superfamily Mesovelioidea: Mesoveliidae Superfamily Hebroidea: Hebridae Superfamily Hydrometroidea: Hydrometridae Superfamily Gerroidea: Hermatobatidae, Veliidae, Gerridae lnfraorder Leptopodomorpha Superfamily Saldoidea: Aepophilidae, Saldidae Superfamily Leptopodoidea: Leptopodidae, Omaniidae lnfraorder Cirnicomorpha Superfamily Joppeicoidea: Joppeicidae Superfamily Tingoidea: Tingidae Superfamily Miroidea: Microphysidae, Miridae Superfamily Cirnicoidea: N~'-idae, Anthocoridae, Cirnicidae, Polyctenidae Superfamily Reduvioidea: Pachynornidae, Reduviidae lnfraorder Pentatomomorpha Superfamily Aradoidea: Aradidae Superfamily Piesmatoidea: Piesmatidae Superfamily Coreoidea: Colobathristidae, Berytidae, Malcidae, Lygaeidae, Largidae, Pyrrhocoridae, Stenocephalidae, Coreidae, Alydidae, Rhopalidae Superfamily Pentatomoidea: Thaumastellidae, Urostylididae, Plataspididae, Cydnidae, Thyreocoridae, Scutelleridae, Dinidoridae, Tessaratornidae, Eumenotidae, Pentatornidae, Acanthosomatidae
Under the heading the valid name and its synonyms are given in chronological order, with reference, information on availability, homonymy, and synonyrnization or change of rank. Information on type-material is given as complete as possible, Category: sex, type-locality; depository. Type material is not given for replacement names and
i.e.
emendations. The category of types is given as follows: HT for holotype, LT for lectotype, ST for syntype, STS for syntypes, ST(S) for unknown number of syntypes (one or more than one), and NT for neotype. For Lectotypes and Neotypes a reference to !heir fixation is given. Names of countries in type-localities are abbreviated as under distributions. Current names for type localities are given between brackets. The depositories used frequently are abbreviated as upper case four character codens in accordance with Arnett & al. (1993). The abbreviations together with full data on !he depositories are listed on the next pages. Depositories not listed are included with their full names. Detailed distributional information (Distr.) is given for each species and subspecies represented in the Palaearctic Region (fig. 1). The names of countries are abbreviated with a unique two letter code as listed on the next page. Russia (RU) is divided in a European and an Asian part, both subdivided in three territories (fig. 2). China is divided in seven territories (fig. 3). Both Kazakhstan and Turkey are divided in a European and an Asian part. The distributional information is listed under four subheadings: Europe (EU), North Africa (NA), Asia (AS) and Extralimital (EL). Extralimital distributions are not abbreviated. New records are indicated with an exclamation mark, whereas doubtful ones are given with a question mark. Untraceable records from the former territories of Czechoslovakia, the Soviet Union and Yugoslavia are given as "CS", "SU", and "YU" respectively. Under references (Ref.) only the most important references are given with an abbreviated indication of their contents between parentheses. In general not more than five references are given for each valid species or subspecies. Additional information is given as notes (Note). Extralimital species included are given between brackets.
• •• .I
Fig. 1. The Palaearctic Region. xi
Abbreviations of depositories
AMNH American Museum of Natural History, New York, N.Y., U.S.A. Museum of Natural History, London, Great Britain Bernice P. Bishop Museum, Honolulu, Hawaii, U.S.A. California Academy of Sciences, San Francisco, California, U.S.A. Canadian National Collection of Insects, Ottawa, Canada Natural History Museum, Beijing, P.R. China Museum of Zoology, Cambridge University, Cambridge, Great Britain Deutsches Entomologisches lnstitut, Eberswalde, Germany EAMF Ecole d' Agriculture, Montpellier, France Entomological Institute, Hokkaido University, Sapporo, Japan EIHU EMAU Ernst Moritz Amdt Universitiit, Greifswald, Germany Ege University, lzmir, Turkey EUIT Gymnasium der Franziskaner, Bozen, Italy GFBI HMUG Hunterian Museum, University of Glasgow, Glasgow, Great Britain HNHM Hungarian Natural History Museum, Budapest, Hungary Institute of Entomology, Czech Academy of Sciences, Ceske Budejovice, IECA Czech Republic Istituto di Entomologia "Guido Grandi", Bologna, Italy IEGG lnstituto Municipale de Ciencias Naturales, Barcelona, Spain IMBS Inner Mongolian Teachers' University, Department of Biology, Huhehot, Inner IMTU Mongolia, P.R. China lnstituto Nazionale di Entomologia, Rome, Italy INER Institut Royal des Sciences Naturelles de Belgique, Brussels, Belgium IS NB Iowa State University Insect Collection, Ames, Iowa, U.S.A. ISUI Institute of Systematic Zoology, Polish Academy of Sciences, Krakow, Poland ISZP Institute of Zoology, Academia Sinica, Beijing, P.R. China IZAS Institute of Zoology and Botany, Tartu, Estonia IZBE Kyushu University, Entomological Collection, Fukuoka, Japan KUEC KUKC Kyungpook University, Taegu, South Korea Charles University, Department of Systematic Zoology, Prague, Czech Republic KUPC LNMD Landesmuseumfiir Naturkunde, Miinster, Germany Linnean Society, London, Great Britain LSUK LUWC Landbovwuniversiteit, Department of Entomology, Wageningen, The Netherlands Museu Bocage, Laboratorio Zoologico, Faculdade de Ciencias, Lis boa, Portugal LZLP MACN Museo Argentino de Ciencias Naturales, Buenos Aires, Argentina MCSN Museo Civico di Storia Naturale "Giacomo Doria •, Genoa, Italy MCNV Museo Civico di Storia Naturale, Venice, Italy MCZC Museum of Comparative Zoology, Harvard University, Cambridge, Massachusetts, U.S.A. MCZR Museo Civico di Zoologia, Roma, Italy MGAB Muzeul de Istoria Naturala "Grigore Antipa", Bucharest, Romania MGFf Museum G. Frey, Tutzing, Germany MHNG Museum d'Histoire Naturelle, Geneva, Switzerland MHNL Musee Guimet d'Histoire Naturelle, Lyon, France Museo Istituto di Zoologia, Universita di Torino, Torino, Italy MIZT MLUH Martin Luther Universitiit, Halle, Germany MMBC Moravian Museum, Brno, Czech Republic MNHN Museum National d'Histoire Naturelle, Paris, France MNMS Museo Nacional de Ciencias Naturales, Madrid, Spain MRAC Musee Royal de I' Afrique Centrale, Tervuren, Belgium MRSN Museo Regionale di Scienze Naturali, Torino, Italy MSIE Shanghai Institute of Entomology, Shanghai, P.R. China MSNM Museo Civico di Storia Naturale, Milan, Italy BMNH BPBM CASC CNCI CNHP CUMZ DEIC
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Abbreviations of depositories, continued MSNT
Museo Civico di Storia Narurale, Trieste, Italy
MVMA
Museum of Victoria, Abbotsford, Australia
MWNH Museum Wiesbaden, Narurwissenschaftliche Sammlung, Wiesbaden, Germany Museo de Zoologia, Barcelona, Spain Zoological Museum, University of Helsinki, Finland Musee Zoologique, Lausanne, Switzerland Museum of Zoology, Lund University, Lund, Sweden National Chung Hsing University, Taichung, Taiwan Narurhistorisches Museum, Base!, Switzerland National Science Museum (Narural History), Tokyo, Japan Narurhistorisches Museum Wien, Austria Narurhistoriska Riksmuseet, Stockholm, Sweden NKUM Nankai University, Deparnnent of Biology, Tianjin, P.R. China NMBA Narurhistorisches Museum der Benediktiner-Abtei, Admont, Austria National Museum of Ireland, Dublin, Ireland NMID NMPC National Museum of Narural History, Deparnnent of Entomology, Prague, Czech Republic National Taiwan University, Taipei, Taiwan NTUC NWAU Northwestern Agriculrural University, Deparnnent of Plant Protection,Yangling, Shaanxi, P.R. China National Zoological Survey of India, Calcutta NZSI OLML Oberosterreichisches Landesmuseum, Linz, Austria OMNH Osaka Museum of Narural History, Osaka, Japan OXUM Oxford University Museum, Hope Entomological Collections, Oxford, Great Britain PMAG Perth Museum and Art Gallery, Perth, Great Britain PMNH Peabody Museum, New Haven, Connecticut, U.S.A. Phyletisches Museum, Jena, Germany PMJG RMNH Nationaal Naruurhistorisch Museum, Leiden, The Netherlands Snow Entomological Museum, University of Kansas, Lawrence, Kansas, U.S.A. SEMC Forschungsinstirut und Narurmuseum Senckenberg, Frankfurt am Main, Germany SMFD Staatliches Museum fiir Narurkunde, Stuttgart, Germany SMNS SNMB Staatliches Naturhistorisches Museum, Braunschweig, Germany SNMC Slovenske Narodne Muzeum, Bratislava, Slovakia TMNH Tianjin Museum of Natural History, Tianjin, P.R. China Institute of Zoology, Ukrainian Academy of Sciences, Kiev, Ukraine UASK uccu University of Kharkov, Kharkov, Ukraine USNM United States National Museum of Narural History, Washington, D.C., U.S.A. UWCP University of Wroclaw, Deparnnent of Entomology, Wroclaw, Poland Universitets Zoologiska Instirut, Uppsala, Sweden UZIU UZMT Zoological Museum, University of Turku, Finland Zoological Institute of the Bulgarian Academy of Sciences, Sofia, Bulgaria ZISB Istituto di Zoologia, Universita di Napoli, Napoli, Italy ZIUN Istituto di Zoologia, Universita di Napoli, Portici, Italy ZIUP ZMAN Zoologisch Museum, University of Amsterdam, Amsterdam, The Netherlands Zoological Institute, Russian Academy of Sciences, St. Petersburg, Russia ZMAS ZMHB Zoologisches Museum, Humboldt Universitiit, Berlin, Germany ZMPA Zoological Instirute, Polish Academy of Sciences, Warszawa, Poland ZMUC Zoological Museum, University of Copenhagen, Copenhagen, Denmark ZMUH Zoologisches Museum, Universitiit Hamburg, Germany ZMUL Zoological Museum, Lvov University, Lvov, Ukraine ZMUM Zoological Museum, University of Moscow, Moscow, Russia ZSMC Zoologische Staatssammlung, Miinchen, Germany MZBS MZHF MZLS MZLU NCHU NHMB NSMT NHMW NHRS
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::..\
ZMAS.
Distr. AS: IN.
Systelloderes utukhengal Linnavuori, 1984 Systelloderes utukhengal Linnavuori, 1984: 19. HT:
o, IQ,
Baghdad; AMNH.
Distr. AS: IQ.
Systelloderes uvarovi Stys, 1970 Syste/loderes uvarovi Stys, 1970c: 438, 440. HT:
o, IQ, Kurdistan, Ugla post; BMNH.
Distr. AS: IQ.
s
lnfraorder DIPSOCOROMORPHA I.M. Kerzhner The very small plesiomorphic bugs included in this infraorder were in many publications placed in one family, named Cryptostemmatidae, Dipsocoridae, or Ceratocombidae. The bugs are living in leaf litter, in decaying wood, in mosses, and under stones at riverbanks. The species diversity is much higher in tropical regions, but some species occur in temperate and cold areas. Representatives of the group are regarded generalized predators of small arthropods, but observations on the feeding are extremely scarce. Th'! most important publications on the group are Reuter, 1891a (first monograph), McAtee & Malloch, 1925 (first comprehensive paper on American fauna with a key to genera), Wygodzinsky, 1950a, 1953 (Afrotropical fauna), Emsley, 1969 (Schizopteridae of Trinidad, with a catalogue of the world fauna), Stys, 1970d (higher classification), 1983 (comparative morphology), 1990 (account on the West Palaearctic fauna). Extralimital families: Hypsipterygidae Drake, 1961, Palaeotropical; Stemmocryptidae Stys, 1983, New Guinea. Infraorder DIPSOCOROMORPHA Miyamoto, 1961 Family CERATOCOMBIDAE Fieber, 1860 A small family, distributed worlwide. About 10 genera and 50 species are described, but the number of undescribed tropical species is very high. All palaearctic representatives belong to one genus, which includes most species and has the largest distribution within the family. Most of the palaearctic species are living in moderately humid leaf litter in forests and meadows, but Ceratocombus corticalis is living under loose bark and in rotten wood of coniferous trees in primeval forests. The palaearctic species overwinter in the egg stage. Family CERATOCOMBIDAE Fieber, 1860 CERATOCOMBIDAE Fieber, 1860a: 267. Type genus: Ceratocombus Signoret, 1852. Extralimital subfamily: Trichotonanninae Stys, 1970, Palaeotropical Region. Subfamily CERATOCOMBINAE Fieber, 1860 Extralimital tribe: Issidomimini Stys, 1970, Palaeotropical Region. Tribe CERATOCOMBINI Fieber, 1860 Genus Ceratocombus Signoret, 1852 Ceratocombus Signoret, 1852: 542. Type species by monotypy: Astemma mulsanti Signoret, 1852 (= Anthocoris? coleoptrata Zetterstedt, 1819). Uchenobia Baerensprung, 1857: 165 (syn. Dohrn, 1858: 229). Type specie-s by monotypy: Lichenobiaferruginea Baerensprung, 1857 ( = Anthocoris? coleoptrata Zetterstedt, 1819). Extralimital subgenus: Tagalonannus Poppius, 1915, Oriental Region. Note: Stys (1990: 8) indicated that he has seen unidentified specimens of this genus from Cl, NW Africa, LE and SY.
Subgenus Ceratocombus Signoret, 1852
Ceratocombus coleoptratus (Zetterstedt, 1819) Anthocoris? coleoptrata Zetterstedt, 1819: 74. STS: SV, parish Uirketorp in Ostrogothia [= Ostergotland] and "villa Abusa" in Scania [ = Skane); MZLU? Bryocoris muscorum Fallen, 1829: 153. New name for Anthocoris? coleoptrata Zetterstedt
6
because of secondary homonymy with Salda coleoptrata Fallen, 1807 (now in Myrmedobia) when both were placed in Bryocoris. Astemma mulsanti Signoret, 1852: 541 (syn. Dohrn, 1858: 229). STS: o. ~, FR. Paris ("Bois des Vincennes"); NHMW? Lichenobiaferruginea Baerensprung, 1857: 167 (syn. Baerensprung, 1858: 189). STS: o, '?, GE, Berlin and PL, Wollin [ = Wolin] Is.; ZMHB? Distr. EU: AU BE BU BY! CZ DE EN FI FR GB GE GR HU IR IT MC NL NR PL RO RU (CT NT ST) SK SP SV SZ UK. AS: IS MG RU (ES FE) SY. Ref.: Jordan, 1941; Kerzhner, 1974; Linnavuori, 1951a, 1958; Melber & Kohler, 1992 (bioi., morph.); Stys, 1959 (nymph}, 1990 (bioi.). Subgenus Xylonannus Reuter, 1891 Xylonannus Reuter, 1891a: 8 (as subgenus of Ceratocombus). Type species by subsequent designation (Oshanin, 1912: 85): Ceratocombus corticalis Reuter, 1889. Ceratocombus alticallus Ren & Yang, 1991 Ceratocombus alticallus Ren & Yang, 1991a: 333,336. HT:
