The types of Recent and certain fossil opisthobranch ...

·, The types of Recent and certain fossil opisthobranch molluscs in the Muséum nalional d'Histoire naturelle, Paris Ángel VAlDÉS & Virginle HÉROS Laboratolre de BiOlogia des Invertébrés Marlns el Malacologie , Muséum naliena! d'Hisloire naturelle, 55 rue de Buffon, F·75231 París ce de>: 05 (France) malaco@mnhn. fr avaldes@casma¡tcalacademyorg

Vaki.a A. & HOros V. 19S8. - ThIJ lyp&a 01 Recen! and certain Io~ opillflobrandl molWJ SCS In \he Muséum natkmal d'Hlslolre nóllUrell., París. Zaosyst.mQ 20 (4): e95·742,

KEYW9RDS typ~ ~~Cl1nlnS,

op isthObr:l llchs.

MnlluSCI, MNHN.

l'nri~.

An51'RJ\ r Thr\lt.l hLJ,/ldl\;!U tillO ti(1)' NC .....'n lou: nr k".·U IU .. mi C>Il t tn~ l _ mlolSil npilfl\ú b ro.n~ d i muUutC ,)tpc'''pccbncn ~ ~jt('d in ,he I...:abcn:lil4,mt' d ~ lnvcn&rb Mpl f1 ~ ~'( MnltlOJWl:.or Ul( Mu.~IIJn.natl \)I\at J¡ liP l:Pl~ "RtIIl'dle (MNHN) Alt! OlIil)ogW:¡j bJ' oti¡;lnnl billl) l)l ~ uml tll1U.ntcd 1l1p l1il b ~dcd ly .....·jmm (.llr'UIill",c
che museum.

Toussellae. Ringicu/A rOllJself4t Rosso et Saubade, 1985: 303, 304, fig. 2. Type locality: 14°22'40"'N - I 7"09'W, off Senegal. HOLa· TYPE: said to be dcposiccd in MNHN in the original publication, but never prcsented to che museum sal/una. RingicuM raBel/1M Morler, 1880: 153. 154. Type locality: Cap Breran Canyon, Bay oE Biscay. SYNTYPE (probably): 1 sheJl, Icg. de Folin (Fig. 3 8 ).

1975b: 349, 350, fig. 14. Type locali[y: Expedition, sen 251, 47°38'N " 08°56'W, Bane de la ChapeJle, 3360-3600 m. HOLOTVPE (by original d~ l­ gnacion): shell (Fig. 4F). crossei. Cylich~1Il crouri Bucquoy, Daucr.enberg et Dollfu.Iype loeaJiry: Walda Expedition, stn CY15, 22"53'5 - 11°56'E, Sourheasn: rn Adamic, 17'56 m. HOLOTYPE (by original designa¡jan, figured by BOLlchet 1975b, 1'1, 4, fig. B): shdl (Fig. 4 E) . Fall1ily SMARAGDlNE1,1,lDAF Thiele, 1925

glauca. Bulla g/auca QllOY et Gaill1ard., 1833:

7"8

352, 353, pI. 26, figs 10-12. Type locali[}'; "Havre: Cancrct", New Ird and, Bismarck Archipebgo. HOLOTYPE (by monoo/py, figured by Ql10y & Gaill1ard 1833, pi. 26, figs 10-12)' specimen, leg. Quoyand Gaimard. Remarru: In acring as fifst reviso] in the meaning of the Anide 24 (ICZN 1(85), PruvotFol (1934b) dc[ermined thar Bulla viridis Rang in Quoy & Gaimard 1833 (placed in the gcnlls Srnaragdinella A. Adams, 1848) has preee:dcnce ove:r its sllJenive synonym Bu/la glaltca Quoy ct Gaimard, 1833. lutea. Bulla lutea Quoy et Gaimarcl, 1833.369, 370, pI. 26, Eigs 40-44. '1ype locality: "Pon DOley" (ee Manokwari), Irian .laya, Indonesia. SYNTYPES: 4 specimens, leg, Quoy and Gaimard. vil·idis. Bulld viridis Rang in QllOY & Gaimard 1833: 350·352, pI. 26, Eigs 13-16. -Iype locality: "Rade d'HlJmata" (" Ul1latae), Gualll (PaciEie Oeean), SYNTYPE,.'): 22 spwlllens (6 uf thelll dissecled) and .? shells, leg. QllOY and Gailllard_ RemarlLAOOHRANCHIDAI:. J.

E. Gray, 1840

sclwammi. TridaclJia schnm¡mi March. 1863a: 41. Type !ocaliry: Guadeloupe (Caribbe:m

Sc:a) SYNTYPES: 4 speclmens. Remada: Deshaycs (1857) esrablished the llC"W gellus Trjdachia Deshayes, J 857, bm did not name a specleo. Mórch (J 863,,) imroduced for che firs[ time che binorninal name Tridachia schrammi in refelence lO Deshayes' dcscription. T schrammj, cum::ntly corlSldcred a synonym

ZOOSVSTEM" - 1996 • 20

r-'J

bicincta. Bulla hicincta Quoy ttGaimard, 1833: 355, 356, pI. 26, figs 31, 32. Type localiry: "Pon du Roi~Geotges" ("" KiJlg Gc::orgc::s Sound), Wesrern Ausrralia SYNTYPES: 2 sheUs gtued ro cardboard, scvetdy bcoken, lego Quoyand Gai mard. elegmu. Aura ekgam Locard, 1886: 535. Type localiry: Quiberon, Franee. SYNTYPES: 4 shells (3 proken) (Fig. 9f). Remarks: According ro Lc::rnche (1948), thi~ is a junior slIbjective synonyrn of Akera bu/lata (Muller,

1776). pirata. Acem spirata 5laad{ in Cossmann & Pissarro 19 13. pI. 55, flg. 244-2. Type localiry: Chcnay, Fronce (late: Paleocene). HOLOTYPE (by mOllorypy, figured by Cossmann & Pissarro 1913 , pI. 55, fig . 244-2): shell (Fig, 9)). Family A1'LVSIIDAE Lamarck. 1809 alba. AplysieUa graviui var alba Vayssiere,

71 1

-

\

~dáA

6:.H...,.V.

190Gb: 67, 68, pi 4, figs 66-68. Type localiry: Obock, Djibouti_ HOLOTYPE (by monotypy, figured by Vayssihe 1906b, pi. 4, figs 66-68): spc:cimel\ (dissccred) + sorne p 3 1t:!i prepared fo r SEM. Remarks: According ro M:uríncz (1996), chis subspecific-rank taxoll is (l. junior synonym of P(laJifcra p(taJí/ud (Rang, 1828). llscifera. Aplysia auiftra Rang, 1828b: 5 1, 52, pI. 4, figs 7-9. Type locality: SaJOe Jean, úyel1J1e, Prench G lI i:l.na. SYNTYPES: 2 specimens, leg_ Richard.. Rcmarks: Accol"ding tu Engd & Hummdinck (1936), this is a junior synonym of Do/abri/ml dolabrifora (Rang, 1828). brasilia1Ja. ApLysia brtuiliana R..1Ilg, 1828b: 55, 56, pI. 8, figs 1-3. Typc locaHry: Brazil. SYNTYPES: 3 spedmens (2 of (hem d isse
260.26 1, pI. 18, Eig.~ 5-7. Type locality: N ew Guinea, Indom~"'¡a. SYNTYFE: 1 specimen (dissected). Remarks: Pruyot-Fol (I 934b) placed chis llpec.ies in the genus Plarydoris Bergh .

1877. lamr'nea. Pldrydoris laminea Risbec, 1928: 86, 87. Hg. 14, pL 1. fig. 11. Typl: locality: Ncw CaJedonia. SYNTYPE5: 2 specimens. maculata. Piatydoris maculatn Bouc.hec, 1977: 3741 , figs 6-8, pI. 2, fi.gs B, C. Type locality: T/¡alassd ( 1970) Expcdi tio n , srn W4 13, 43°50N - 06~ 09W, Bay of Biscay, 500-540 ffi _

ZOOSYSTEIdA • 1998 • 20 (~I

Family MIAMlRlDAI! Bergh, 1891 rugosa. Gravieria rugosa Vayssiere, 191 2: 30-32, pI. 10, figs 144 -1 49. Typc: locaJi cy: - 103. Typt loU~Ji ry: R:cpl"Jlin ha, s.;o 1~m~. \,'{Ir::s, Afl'lCl, HO u)Ty]l r-~ (by urlgmíl) dl:iligl1:u,uu. figurcd by Orcea et al. 1996, flgs 120, 100B. f'Jt:Ii

",l

10lB); specimca, lego RoJán and Fernandcs. Flmily DIlNDRODorUDlDAE O'Oonoghuc, 1924

Glon¡J//,rlJ sy'1I!/fC~~~ RnJIIl.ln , 1990; 27j-'r1? fi\l' le . 60 , SA, ~ I) (la ,..,,). angolemis, Dendrodoris ango/enfiJ Valdés et Ortea in Valdés et al. 1996: 20-23, fj gs 2F, 11 , 1aA·D. 11. 11.. 1ypc loc ~ !.[y. SI. GiIlts ltoof.

q m1lldr,',·uJ.

~ Riunion ()¡.di:¡n OC
1 par.llype. catena, N.ryllidia catma Pruyot-Po\' 195Gb: 70-72, figs 6, 7. Type localicy: Mayutte ¡tnd Mauridus (lndian Oce:an). SYNTYPES [one figurcd by Pruvot-Fol 195 6b, figs 6 (m parc), 7] : 2 specimens, leg, MathieUi 1 specimen (di ~ ­ sected), lego Cloué. Remarks: Accordmg ro Bnlllckho!St (1993), chiS is ,1 junior ~ynonym of Phyllididla uyla,¡ica (Kela~rt, 18 59). dlUJtullbergi. Phyliidia dautunb~rgi Vay.ssihe, 191 2: 85· 87 , pI. 1, figs 14 , 15. Type loealiry' "Nord d'Ambouli ", Golrc de TadjollJ'a, l1jiooUlj, HOLO'I'Y}JI!. (by rnnnD'YPY, flgul-ctl

Drunckhom & WiUan (I989), rrus is 11 junior syllonym o f Phyllidjajlava Aradas, 1847. bayi. Fryeria bayi BOllcher, 1983: 65-68, Hgs 1, 2. Type localiry: Puma Blanca. Corsica (Mediterrallean Se-J.). HOLOTYPE (by origiI..y Vllrt-'1i~~ 1912, pL J. I ¡~ 14. 15): . ptcinal design ation, fl gured by Boucher 1983, meD, let, C fJ.vlc!, R~mub! Thc: munr PhyJtii/¡rl '¿IJItM'lÚCl'gi WilS flDl ly illrn'!ducN by fig. 1): l'IJQCimcli 1'- I pillllrype.. V.fl)'SS.t:fC O9ll) wirherll l dcscripfjOI1 {lUlr7fm btlrtlli. PI"ltidiDpliJ (¡rrf.bi V:lPlihe. 19Q 2~ muhn"j. ürundmllUt (199'3) pl"cecl thi~ /o'Peo231·242. pL 9. foil' 10· 16. pI. w. ¡¡g. 17. Typ, loctdit),: ~iJiJIJt.U' 0883~ EXJ)fi{ jlioft , u n l4 1. cíes In j he~llwPl"IJM¡tlptlJ IkIEh. 187:5. -4'j" ~ 9,OO'N .. n4 ~ t)9 ,4 tl ' W, lb)' tlr Bku;:tty, fotlltl . rbjllitlJIl fl{/!{I Aad..u., 1S47! 121 . Type 1480 lll. J-IOLOTYPE (by monotypy, figured ¡oCllity. Xlc-ndi , Gcl'KI. MIlIc.. NP.on'Pl:.. by Vayssicre 1902. pis 9 . 10 :md Valclts & (tlt:llgl1:ltl-d by HtunckboDt & W iJb n 1989J Om:a 1996, fi g. l B): specimen (dlS5eCroo) . ~Ó9, fgwM by 8nm,;;klu:lI'iIf 6:. WBhm 19R~, borhonica, PhyiLidlfl borbonica unavailable. fig. 1): spcclmen + phom. Remada. Brunckhofllf (1 993) studied one gof4.1i. Reticulidia gofasi Valdés el Ortea, 1996: 7, 8, figs .H , 4C, 5 (in pan), 8. Type loca· ~ p e cimen d eposited in MNHN labelled "Phytlidia borbonica Cuv." , and pointed ou r lit y: Seam ount 1 Expeditioil, stn DW61,

7

7'2

1

36°40.02'N - 14°16.00'W, Josephine Baok, Nonh Eastern Atlantic, 200-205 m. HOLOTYPE (by original designation, figllred by Valdés & Ortea 1996, fig. IF): spccimen + 2 paratypes. gynenoplA. Phyllidiopsis gym:nopla BOlicher, 1977: 50-53, figs 18, 19, pI. 2, fig. E. Type localiry: Bim;ores Expedition, sto 159, 3r26'N - 25°51'W, off Santa Maria. Azores. 525-600 m. HOLOTYFE (by original dcsignation, figured by Boucher 1977, figs 18. 19. pI. 2, fig. E and Valdés & Onea 1996, fig. le): specimen (dissected). Remarks: According to Vald¿s & Orrea (1996), rhis 1S a junior synonym of Phyllidiopsis berghi Vayssiere, 1902. honloni. PhylLidia honlom Risbec, 1956: 22, 23, pI. 14, figs 71-75, pI. 15, flgs 79-8 1. Type lacality: Han Lon, Nha Trang, Vietnam. LECTOTYPE (sdected by Brunckhorst, 1993): specimen + 3 paralecmtypes. Remada: According te Brunckhorst (1993), chis is a junior syoonym of Phy¡¡idia varicosa Lamarck. 180l. krempfi. Phyltidiopsis krempfi Pruvoc-Fol, 1957: 120, 121. figs 41-49, pI. 1, flgs 7, 8. Type localit y: Nha Trang, Vietnam. HOLOTYPE (by monotypy, figured by Pntvor-Fol 1957, figs 41~49, pI. 1, figs 7, 8): specimen. nigra. Phyltidia nigra van Hasselt, 1824: 244. Type localiry: Cu bo Hon "1Je Poulo Cecir de Mer", Vietnam. NEOTYPE (designated by Brullckhorsr, 1993: 55): specimen. Remadts: Brunckhorsr (1993) placed chis speeies in ,he genus PhyllididÚt Bergh, 1869. ocellata. Phyllidia oceilata ClIvier, 1804b: 269, pI. 18, fig. 7. Type locality: Timor, Indonesia. LECfOTYPE (sc:lo:: eto:: d by Brunckhorst 1993. figllrd by Pruvot-FoI1956b, fig. 2); spedmen (dissected), lego Péron and Lo:: sw:ur. pu/itu";. Phyliidia pulitzeri Pruvoc-Fol, 1963: 566-569, figs 1-6, pI. 1. Type locality: Portonno, Gc:nova, Italy. HOLOTYPE (by original designaríon, figured by Pruvot-FoI1963, ngs 1-6, pI. 1): speclmen, lego Pulitzee. NEOTYPE (designared by Wagele, 1985: 65. figUfCd by Brunckhom & Willan 1989. fig. 1): spedmen + phoro from Xlendi, Gozo, Malta. Remarks: Wagele (1985) designated a neotype of Ihis species because at thu IÍme me holory-

pe was considered lose. Laree, Brunckhorst & Willan (1989) selected [he same specimen as neorype of Phyllidia flava Aradas, 1847, to fu: rhe synonymy berween borh nominal species. However, we have rediscovered the holorype of Phyllidia pulitzeri in MNHN, and therefore this case should be submincd ro the Commission (ICZN 1985: Artide 75h). pustulosa. Phyllidia pustulosa Cuvier, 1804b: 268, pI. 18, fig. 8. Type localiry: Timar, Indonesia. LECTOTYPE (se!ecred by Brunckhorst 1993, figured by Cuviel 1804b, pI. A, fig_ 8 and Brunckhorst 1993, fig. 27A): specimen, lego Péron and LeSlleur. Remarks; This is rhe rype species of the genus Phy¡¡idi~Lla Bergh. 1869, by subsequem designatian by Brunddlorsr (1993). serenei. Phy¡lidia serenú Risbec, 1956: 24, 25, pI. 16, 82~84, pI. 17, figs 86-89. Type locality: "fle Poulo Cecil' de Mer" (= Cu Lao Hon), Vietnam. LECfOTYPE (sdecred by Brunckhors[, 1993): specimen + 3 paralectotypes. Remarks: Brunckhorsc (1993) designatcd che lecrocype of Phyllidia smmd as Ileocype of Phyltidúl!a nigra (van Hasselt, 1824), and [herefore both mImes become objecrive synonyms. trili!ledta. Phyltidia tr¡lineata Cuvier, 1804b: 268, pI. 18. figs 1-6. Type locali[y: La Réunion, Jndian Oc(': an. SYNTYPE (figured by Cuvier 1804b, pI. 18, figs 1-4): 1 specimen (dissecred) (Fig, lOB). NEOTYPE (designaced by BrunckhoJ'st 1993: 27, figured by Brunckhorst 1993, flg. 23): specirncn, lego Drivas and Jay. Remarks: ClIvier (1797) established the generic name Phyll¡dia far a single specimen collo::cred from "tlo:: Bourbon" (_ La Réunion), but ¿id not name rhe specics. Lamarck (I801) erected the specific name Phyllidia varicosa for ,he specimen seen by Cuvier (1797). Cuvicr (1804b) commented rhar ,he species on which he himsd f based rhe genus f'hyllidia must be named Phyltidia trilincata insrcad of P varicosa because che former name is more appropriate. At the same nme, he describd and figured P. trilineata based on the original specimen (from b Réunion) and rwo additional individL1als (considerably smaller io size) collecred by Péron from "Mer des Indes". The examinaríon of me specimen

ngs

ZOOSYSTEMA • HI9!l • 20 (4)

• OpiSlbobJ;l\Ch mcUw< IJIP< TYrE: I ~p mmen qu.oyi, NUDlBRANCHIA-AEOLlDINA Family FlABELLINIDAE Bergh, 1889

baetiea. Flabdlino bouica Garda-Gómez, 1984: 61..{í4, pl.s 1, lA Type locrrlity: T:lrifa, Spa in. HOLOTYPE (by original designatíon); speci-

n~

lI;¡ctlIÍf¡'~

19_ T)'pI:

1mT)'" Pruml-Fol. 1934b: 56.

Lm.~l j ry;

un1":t1own. H OlOTYPI1

(by lIIoaotypy): IIpcclmc::n (Ji~!"ói!:cltd) . I C!~

Quoy Ilnd G:lim.ud. rnn.ktJII;. Awlülir'llJi, lIf'm1rri JJr·u\'ot-Fol. 1956:r..: ;228-231, 6~s 1- 11 TyP'! loctL¡p IJc K"(lkur:r., liJ.:lffiOru (Pltd flc. CIc.eu.r¡), SYNTYPE! 1 !JM:umen (dissccred), leg. Ranson. Remacks: This is the type species of [he genus At(}lidiopsis

meno

Pruvoc-Fol, 1956, by onginal designadon. IJdlVNq¿ Vil'JIJJfllo bnr.--n'u VAY'liicre. I 1.3= 2~ 'Sy t-ií. Typc I ~icy.: m~u n i l liT ri:ve.I· I ~d~\~. 8c:1Ulh',... Boty. SW COOllt ~lf Famlly GlAUCIDAf. J. E. Gray. 1827 Nu~~r,1 ~ rnl ra , Run¡". HO L.OTYPE (hr monoryp)'. fI~Ufc..l by y~ y!{tl ~i: 19 1~, fi~ 1·6): II.fhn.. )JWtn¡¡".. dOIA G~ ,d u - .. ama: l't Ct.-t'l'Ct';r. $pccimen (dlssccred) I?!}O: 73'1-740. 6~i 1 ~9. 1 pdoClUry: [laia tr ¡JI!)'~ , HiiuJ r~

py):

Iillnl rt.lk. AnIMII\1t

M. l. c. L Jc.

longicouda Quoy et Gaimard, 1832: 288·290, pI. 21, 'figs 19,20. Type 10,",lity: Cook Sn;1it. New ualand. SYNTYPES: 6 spcdmens. Remarks: ThlS is rhe type spccies of che genus Doficheolis Finlay, 1926, by monmypy.

i"

lY\SI'J.

O{/lrnNW\."u 1'1 Jd ~d,.","M If~r ¡tltl

i"" folm

,Utltit;,; IJ'lI

,~~ uuJu1

f'!,yMJ&m. ir grami,

Jo'

jrt

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'r"(II!JI'lC 1 (1) . tnlprln)L!I'¡~ ¡mp~'

rúl~, Pul"ll. llnulI K. 19)7.-0p¡nMbr,J "~ h t..or l t.P1n 1..}ow7Utl uf 1(1'" l?tjUQ~fiI /lf ¡f~llmf k)wv/JJ Imlxl'uJ

ÚrtUJmtIJ' 'j I 289-344

Rnflelrcrol M .. U~ I!. &: 01101 J. 1 1}~4. - 11,c,. i¡jóll ele 1,.. f) odd4CCJ. ( .Mu lhu~i O\li.¡lwbu.ndib) dLi A¡}AnDCD Nonbu wlrl LJIII ~

:l cuu~.P~qo

m>rcWiI·

urJhw1ir. &{I.t'ttTlliJ /';{nL-I~t'~ :t01 4.(.7-21'/.

l\tlIXl)t'! E 1975.1, Admowledgements This paper would have not been possible wirhom the illvaluable help of S!:rge Gafas dming me inv!:IHory of me t)'pe coll!:cuon daoor;1tion

ana

732

t'N

1:i.~f!~ /,~lft(rt.'ll l'rxpt.bil"" IÚ' t;,mlk f'N/lfl/iu"

Fam.ly FJONJDAE J. E. Gray. 1857

longicauda. Eolidia

¡ 11~ D"dtrél ; 7~ S6,

(e~I.) . l>tm'~UÑIII & rE,x",pff. 11" rr..1¡nl

NuiJitu;uxlM.",

110UVl!ll~IJ.

411;11

córes du Sén4;al. Vif (1 Milicll (A) 25 . 1 L9- 132.

1975b. - Opi$lhobrandle.~ de profondeur de I'Ocba Adantlque. 1. - Cepha.laspidea. Cahim de Biawgir Marine 16 : 3 17-}65. - 1977. - Opisthobranches de profondenr de -

ZOOS'I'STEMA • 1998 • a ~ (4)

[

•

Opamooruteh mol' ..", 'YP"