Zootaxa, Review of the Australian genus ...

Zootaxa 2351: 39–48 (2010) www.mapress.com / zootaxa/

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Copyright © 2010 · Magnolia Press

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Review of the Australian genus Desudaboides Musgrave with descriptions of four new species (Hemiptera: Fulgoromorpha: Fulgoridae) JEROME CONSTANT Royal Belgian Institute of Natural Sciences, Department of Entomology, Vautier street 29, B-1000 Brussels, Belgium. E-mail: [email protected]

Abstract Four new species of Desudaboides Musgrave, 1927, e.g. D. fletcheri n. sp., D. melindae n. sp., D. nigrifrons n. sp. and D. rentzi n. sp. are described from Australia. A key to the species, distribution maps and colour illustrations of the five species of Desudaboides are given. Male genitalia are illustrated for D. fuscomaculata Musgrave, 1927, D. melindae and D. nigrifrons. Key words: Lantern-fly, Northern Territory, New South Wales, Queensland, Fulgoroidea

Introduction The Australian genus Desudaboides Musgrave, 1927 was erected for a species from Queensland, Desudaboides fuscomaculata Musgrave, 1927. Musgrave (1927) believed that the genus is close to Desudaba Walker, 1858, a view followed by Metcalf (1947). Lallemand (1963) did not share this view and placed it in the Poiocerinae: Poiocerini: Poiocerina, close to the Oriental Polydictya Guérin-Méneville, 1844 and Gebenna Stål, 1863, and the Australian and Papuan Birdantis Stål, 1863, while he placed Desudaba in the Aphaenini of the Aphaeninae. Nagai & Porion (1996) followed these views and illustrated Desudaboides fuscomaculata. This classification is followed here. The present paper describes four new species of this genus which have been identified from Northern Territory, Queensland and New South Wales.

Materials and methods The male genitalia were extracted using a needle blade from softened specimens and boiled in water for further cleaning. They were placed in glycerin for conservation. Distribution maps were produced by the software CFF 2.0 (Barbier & Rasmont, 2000). For the labels of the types, the wording on each single label is limited by square brackets. The type specimens of the new species here described bear a label of the following type: [Holotype / Paratype ♂ / ♀ Genus species n. sp. Jérôme Constant det. 2010]. The following acronyms are used for the measurements (taken as in Constant, 2004): BF, breadth of the frons; BT, breadth of the thorax; BTg, breadth of the tegmina; BV, breadth of the vertex; LF, length of the frons; LM, length of the mesonotum; LP, length of the pronotum; LT, total length; LTg, length of the tegmina; LV, length of the vertex.

Accepted by M. Fletcher: 24 Dec. 2009; published: 4 Feb. 2010

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Acronyms used for the collections (name of the curator in parentheses). AMS ANIC ASCU MNHN NTM RBINS

Australian Museum, Sydney, New South Wales, Australia (M. Moulds) Australian National Insect Collection, CSIRO, Canberra, Australian Capital Territory, Australia (T.A. Weir) Agricultural Scientific Collections Unit, Orange Agricultural Institute, Orange, New South Wales, Australia (M.J. Fletcher) Muséum National d’Histoire Naturelle, Paris, France (T. Bourgoin and A. Soulier) Northern Territory Museum, Darwin, Northern Territory, Australia (G. Dally) Royal Belgian Institute of Natural Sciences, Brussels, Belgium (P. Grootaert)

Taxonomy Family Fulgoridae Duméril, 1820 Subfamily Poiocerinae Haupt, 1929 Tribe Poiocerini Metcalf, 1938 Genus Desudaboides Musgrave, 1927 Type-species: Desudaboides fuscomaculata Musgrave, 1927 by original designation and monotypy. Desudaboides Musgrave, 1927: 47; Metcalf, 1947: 92; Lallemand, 1963: 8; Nagai & Porion, 1996: 15.

The genus is characterized by the following combination of characters: (1) head as broad as pronotum; (2) frons broader than long and with lateral margins sinuate, longer than the clypeus; (3) clypeus broader than long; (4) vertex excavate; (5) mesonotum tricarinate; (6) tegmina at least three times longer than broad, roundly pointed apically; (7) clavus open with vein A1+A2 short and ending in sutural margin; (8) hind wings shorter and broader than tegmina. The genus is endemic in Australia where one other genus of Poiocerinae is represented (in N Queensland): Birdantis Stål, 1863. The latter is easily separated by the head being narrower than the thorax.

Key to the species of Desudaboides 1. 2.

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Hind wings red or orange basally ................................................................................................................................. 2 Hind wings without red or orange marking .................................................................................................................. 3 Tegmina, frons, vertex, pro- and mesonotum strongly marked with black spots (Figs 1I–J); base of hind wings largely red (Fig. 1I) ........................................................................................................................... D. fuscomaculata No black spots: tegmina brown variegated with pale yellow, orange and green (Fig. 1 G); frons red with carinae brown-green (Fig. 1H), vertex and pronotum red; mesonotum red to red-brown; base of hind wings largely orange (Fig. 1G) .......................................................................................................................................................... D. rentzi Abdomen with tergites 5–8 red (Fig. 1A); 2 small black spots on disc of frons (Fig. 1 B) ........................ D. fletcheri Abdominal tergites black, frons differently marked ..................................................................................................... 4 Disc of frons with large black marking (Fig. 1F); base of tegmina rosy (Fig. 1E); size less than 24 mm ..................... .................................................................................................................................................................. D. nigrifrons Disc of frons without black mark (Fig. 1D); base of tegmina brown (Fig. 1C); size more than 25 mm... D. melindae

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FIGURES 1A–J. Desudaboides: habitus in dorsal view and frons in normal view. A–B, D. fletcheri. C–D, D. melindae. E–F, D. nigrifrons. G–H, D. rentzi. I–J, D. fuscomaculata.

FULGORIDAE: DESUDABOIDES

Zootaxa 2351 © 2010 Magnolia Press ·

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Desudaboides fuscomaculata Musgrave Figs 1I–J, 2, 5. Desudaboides fuscomaculata Musgrave, 1927: 47

Etymology. fuscomaculata (adj., Latin): from fuscus, brown, and maculatus, spotted. Literally “spotted with brown”, the name refers to the colouration of the tegmina. Type material examined: Holotype ♂: [Holotype Desudaboides flavomaculata Musgrave ♀] [Chilla, Jan 24] [Male, stated in error as ♀ in paper, Det. by A. Musgrave.] [k55909] [Holotype ♂ Desudaboides fuscomaculata Musgrave, 1927, Jérôme Constant det. 2009] (AMS). Allotype ♀: [Allotype Desudaboides flavomaculata Musgrave ♂] [Chilla, Jan 24] [Female, stated in error as male in paper, Det. by A. Musgrave.] [k55910] [Paratype ♀ Desudaboides fuscomaculata Musgrave, 1927, Jérôme Constant det. 2009] (AMS). Coordinates of Chinchilla (= Chilla): 26°45'S 150°38'E. Notes: The species has been described under the name fuscomaculata (Musgrave, 1927) but both type specimens bear labels indicating flavomaculata. The other labels on the specimens do not allow any doubt that they are the types. Additional labels with the right name have been attached to both specimens to avoid future confusion. Nagai & Porion (1996) provide a habitus illustration of the species but erroneously stated that the type specimens of D. fuscomaculata are deposited in the collections of the Macleay Museum, University of Sydney.

FIGURES 2A–C. Desudaboides fuscomaculata, genitalia ♂. A, pygofer, anal tube and gonostyle, lateral view. B, anal tube, dorsal view. C, pygofer and gonostyli, postero-ventral view. An: anal tube; G: gonostyli; Py: pygofer. Scale 1 mm.

Other material examined: 2 ♂: 40 Mile Scrub near Mt. Garnet, N. Qld, 9.i.1973, M.S. & B.J. Moulds (AMS) coordinates of 40 Mile Scrub: 18°5’S 144°50’E; 1 ♂: 40 Mile Scrub, 65 km SW of Mt. Garnet, N. Qld, 19.xii.1974, M.S. Moulds (AMS); 2 ♂: idem, 15.iii.1982 (ASCU); 2 ♂, 1 ♀: 40 Mile Scrub, 40 mi SW of Mt. Garnet, N. Qld, 19.xii.1974, M.S. Moulds (ASCU); 1 ♀: 40 Miles Scrub, 64 km SW of Ravenshoe, N. Qld, 7.i.1976, mv lamp, D.K. McAlpine (AMS); 2 ♂, 2 ♀: Chillagoe, GPS 300, 11-12.iii.1997, light trap, Th. Bourgoin (1 ♂, 1 ♀: MNHN; 1 ♂, 1 ♀: RBINS) coordinates of Chillagoe: 17°9'S 144°32'E Additional data: Mareeba (Nagai et Porion, 1996). Coordinates of Mareeba: 17°0'S 145°26'E. - specimens in ANIC (pers. comm. Tom Weir, 2008): 1 ex.: Emu Ck, 27 km SW of Dimbulah, QLD, 2526.xi.1981, J. Balderson, coordinates of Emu Creek: 17°20’S 144°57’E; 1 ex.: Lolworth Station, N.

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Queensland, 29.vi.1907, Mrs Black, coordinates of Lolworth Station: 20°11’S 145°1’E; 2 ex.: 40 Mile Scrub, 40 miles SW of Mt Garnet, NQ, 9.i.1973, G.J. Brooks; 4ex.: idem, 26.ii.1974; 1 ex.: 40 Mile Scrub, 55 miles SW by S of Mt Garnet, 8.xii.1985, J. Balderson; 5 ex.: 40 Mile Scrub, 4.x.1989, L. Ring; 3 ex: idem, 12.iii.1994, L.R. Ring; 6 ex.: Mt. Garnet, 14.iv.1944, coordinates of Mt. Garnet: 17°41’S 145°7’E. Diagnosis. The species is immediately recognized by the following combination of characters: (1) frons pale yellow to red with 4 black spots on disc (Fig. 1 J), (2) hind wings with base red (Fig. 1 I), (3) abdominal tergites 3 to 8 bright orange (Fig. 1 I), (4) tegmina with numerous black-brown spots, with base rosy red and with apex hyaline (Fig. 1 I). Genitalia ♂: see Figs. 2A–C. Biology. Nothing is known except that five of the 15 examined specimens have been caught by light trap. Distribution (Fig. 5). Eastern Queensland. The distribution seems to roughly follow the Great Dividing Range.

Desudaboides fletcheri n. sp. Figs. 1A–B, 5. Etymology. fletcheri: dedicated to my friend, Dr Murray J. Fletcher who sent the only known specimen for study and always kindly supported the work of the author. Material examined. Holotype ♀: [East Ballina, N.S.W, 10 March 1964, I. Bathgate, at light] [MJF collection, MJF003192] (ASCU) coordinates of East Ballina: 28°52'S 153°35'E. Diagnosis. The species is immediately recognized by the following combination of characters: (1) frons pale yellow-brown with 2 small black spots at base of disc (Fig. 1 B), (2) hind wings without red or orange (Fig. 1 A), (3) abdominal tergites 5 to 8 red (Fig. 1 A), (4) size less than 24 mm. Description. LT (extrapolated): ♀ (n = 1): 21 mm. Head: pale yellow-brown, with 2 baso-lateral points on disc of frons and frons-vertex joint black (Figs. 1 A–B); median and lateral oblique carinae on disc of frons well marked (Fig. 1 B); pedicel large, inflated and reniform (Fig. 1 B); labium reaching median trochanters; ratio BV/LV = 4.9; BF/LF = 1.5. Thorax: pale yellow-brown, including tegulae; row of 4 black dots on pronotum, behind head; mesonotum with black dots: row of 7 along posterior margin of pronotum, row of 4 on disc and one on scutellum; 2 small black spots on lateral thoracic pleura; mesonotum black (Fig. 1 A); ratio LP+LM/BT = 0.9; LM/LP = 2.2. Tegmina: apical half hyaline with veins pale yellow-green; basal half pale yellow-green with veins brown; costal cell pale orange-brown; all densely covered with rounded, black spots, often confluent, larger ones with central pale spot covered with white waxy secretion (fig. 1 A); ratio LTg/BTg = 3.25. Hind wings: apical half hyaline; basal half black with large, ante-basal, milky white patch; white patch marked with black between veins R and M and along claval joint; veins black, vein Sc white basally (Fig. 1 A). Legs: all legs pale yellow-brown with apex of tarsi and of spines of hind legs black; hind tibiae with 4 lateral and 7 apical spines (Fig. 1 A). Abdomen: dorsally, black with sides of tergite 4 and tergites 5 to 8 red (Fig. 1 A); pale yellow-brown ventrally. Biology. The only known specimen has been caught at light trap. Distribution (Fig. 5). NE New South Wales.

Desudaboides melindae n. sp. Figs. 1C–D, 3, 6. Etymology. melindae: dedicated to Dr Melinda L. Moir who collected four of the six specimens of the type series.

FULGORIDAE: DESUDABOIDES


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Material examined. Holotype ♂: [N.S.W.: Rummery Park Campsite, Nightcap NP; 28.59912°S153.37755°E, 16 Dec 2008, MV light; 363m; M.L. Moir & K.E.C. Brennan] (ASCU); 2 paratypes ♂: [dry rainforest, N.S.W., Broken Head Nat. Res., 28°42’32”S 153°36’52”E, 70m, 15.xii.2008. mv light, M.L.Moir & K. Brennan] (RBINS); paratype ♂ : [NSW, Big Rocky Dam, Nightcap Nat. Pk., 216m, 28°38’15”S 153°20’47”E, 18.xii.2008, mv light, M. Moir & K. Brennan] (ASCU); paratype ♂ : [Maroochydore, Qld, 11 Dec. 1984, G. Theischinger] [MJF collection, MJF003027] (ASCU) coordinates of Maroochydore: 26°39’S 153°06’E; paratype ♀: [Bluewater Range, 25.ii.1989, L. Ring] [MJF collection, MJF0003191] (ASCU) coordinates of Bluewater Range: 18°54’S 145°57’E.

FIGURES 3A–C. Desudaboides melindae, genitalia ♂. A, pygofer, anal tube and gonostyle, lateral view. B, anal tube, dorsal view. C, pygofer and gonostyli, postero-ventral view. An: anal tube; G: gonostyli; Py: pygofer. Scale 1 mm.

Diagnosis. The species is immediately recognized by the following combination of characters: (1) disc of frons without black spots, small spots only on fronto-clypeal joint, (2) hind wings without red or orange, (3) abdominal tergites black, (4) size more than 25 mm. Description. LT: ♂ (n = 5): 27.1 mm (26.6–28.1); ♀ (n = 1): 32.2 mm. Head: pale yellow-olivaceous with minute red spots; black: middle of hind margin of vertex, transverse spot on each side of disc of vertex , frons-vertex joint, small median and small oblique lateral lines along dorsal margin of frons, 4 small spots along fronto-clypeal joint, often 2 small longitudinal stripes on postclypeus (Figs 1 C–D); frons marked with olivaceous before antennae (Fig. 1 D); labium brown, surpassing hind coxae; median carina of frons obsolete; pedicel large, reniform, green-brown (Fig. 1 D); ratio BV/LV = 5.8; BF/LF = 1.3. Thorax: yellow-olivaceous with minute red spots; 2 small red spots on anterior half of pronotum, posterior half infuscate; mesonotum with row of 5 black spots along margin of pronotum, row of 4 black spots on disc, lateral ones sometimes confluent with spots of anterior row; 2 small black spots near hind margin and large black spot at base of scutellum, latter joining median anterior black spot by black line on median carina (Fig. 1 C); small black marks on lateral thoracic pleura; ratio LP+LM/BT = 1.0; LM/LP = 2.4. Tegmina: slightly more than basal half and costal cell pale yellow-brown, apex hyaline; all covered with large, irregular, brown to black markings; hyaline large patch along sutural margin after clavus (Fig. 1 C); ratio LTg/BTg = 3.2. Hind wings: black with apical 2/5 hyaline, often infuscate in males; elongate pale blue-white markings near base, turquoise in females, between veins A1 and A2, between veins M and SCu and along vein Sc; veins black-brown (Fig. 1 C).

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Legs: pale yellow-olivaceous with irregular black longitudinal markings; I and II with tibiae black apically and obsolete black ring at middle, and tarsi black; hind tibiae with 6-7 lateral and 7 apical spines black apically; apex of 3rd hind tarsomere black (Fig. 1 C). Abdomen: tergites black with margin finely bordered with pale yellow (♂) (Fig. 1 C) or pale turquoise (♀); sternites pale yellow-green to pale yellow-brown, with broad basal transverse black marking. Genitalia ♂: see Figs. 3 A–C. Biology. Four out of the six specimens have been caught at light trap. Two specimens have been captured in a dry rainforest. Distribution (Fig. 6). Eastern Queensland and NE New South Wales, not far away from the coast.

Desudaboides nigrifrons n. sp. Figs. 1 E–F, 6. Etymology. nigrifrons: from niger (adj., Latin) = black and frons (Latin) = frons. The name refers to the colour of the frons. Material examined. Holotype ♂: [Kuranda, N. Qld, Speewah, Ganyan Dr., 20.x.2003, P. Erbe. m.v. lamp] [ASCT HE029967] (ASCU) coordinates of Kuranda: 16°49'S 145°39'E. Diagnosis. The species is immediately recognized by the following combination of characters: (1) frons with large black marking on disc, (2) hind wings without red or orange, (3) abdominal tergites black, (4) size less than 24 mm.

FIGURES 4A–C. Desudaboides nigrifrons, genitalia ♂. A, pygofer, anal tube and gonostyle, lateral view. B, anal tube, dorsal view. C, pygofer and gonostyli, postero-ventral view. An: anal tube; G: gonostyli; Py: pygofer. Scale 1 mm.

Description. LT: ♂ (n = 1): 17.6 mm. Head: pale rosy-yellow; vertex with 2 black spots along posterior margin and 3 black spots on anterior carina, slightly extending on frons (Figs. 1 E–F); vertex constricted in middle (Fig. 1 E); disc of frons with large black patch irregularly extending to sides of frons; clypeus with black patches on margins and 3 pairs of brown spots along median line (Fig. 1 F); labium brown apically, surpassing hind coxae; carinae of frons well marked (Fig. 1 F); pedicel large, reniform; ratio BV/LV = 8.1; BF/LF = 1.4. Thorax: pale rosy-yellow; pronotum with 2 small black spots behind vertex and small black spot behind eyes; disc with 2 pairs of irregular black markings; 3 small black spots along hind margin; mesonotum with 6 black spots along pronotum margin, sometimes confluent; disc with angulous black marking on each side, anteriorly joining median black spots; median carina black; scutellum with basal black spot at each side and larger black spot in middle; metanotum black with transverse pale yellow marking (Fig. 1 E); lateral thoracic pleura irregularly marked with black; ratio LP+LM/BT = 0.9; LM/LP = 2.0. FULGORIDAE: DESUDABOIDES


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Tegmina: basal half variegated brown, with base and clavus rosy; rosy areas with black-brown spots; apical half hyaline with irregular dark brown marks and irregular, broad, ante-apical, oblique, brown line; veins rosy-brown to brown (fig. 1 E); ratio LTg/BTg = 3.0. Hind wings: basal half black with elongate white markings between veins A1 and A2, between veins M and SCu, and along vein Sc; apical half hyaline; veins black, Sc white near base (Fig. 1 E). Legs: pale yellow strongly marked with irregular black patches; tibiae I and II with 3 rings and tarsi black; hind tibiae with 4 lateral and 7 apical spines; spines black apically (Fig. 1 E). Abdomen: tergites black (Fig. 1 E); sternites yellow-brown with hind margin paler. Genitalia ♂: see Figs. 4 A–C. Biology. The only known specimen has been caught at light trap. Distribution (Fig. 6). NE Queensland.

FIGURE 5. Distribution map of Desudaboides fuscomaculata, D. fletcheri and D. rentzi in Australia.

Desudaboides rentzi n. sp. Figs. 1G–H, 5. Etymology. rentzi: dedicated to my friend, Dr David Rentz (Kuranda, Queensland) who is one of the collectors of the holotype of the new species. Material examined. Holotype ♀: [NT Lichtfield N.P., Florence Falls, M V, 13°03’S 130°47’E, 29 Oct. 1991, Wells & Rentz] (NTM); paratype ♀: [12.46S 132.39E, 12 km NNW of Mt. Cahill, N.T. 20 May 1973, Upton & McInnes] (ANIC); paratype ♀: [NT, Humpty Doo, 12 March 1994, N. Richardson, at U.V. light] (RBINS) coordinates of Humpty Doo: 12°38'S 131°15'E; paratype ♀: [Beerimah, N.T. via Darwin. 5.xii.1974,

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A. Allwood.] [3754] (NTM) coordinates of Berrimah: 12°25’S 130°55’E; paratype ♀ : [12.44S 130.58E, Darwin River, 16km SW by S of Noonamah, 16.v.1974, N.T., T. Angeles & W. Mollah] [3755] (NTM).

FIGURE 6. Distribution map of Desudaboides melindae and D. nigrifrons in Australia.

Diagnosis. The species is immediately recognized by the following combination of characters: (1) frons red, often with oblique carinae olivaceous, (2) hind wings with bright orange base, (3) abdominal tergites 3 to 8 red, (4) tegmina opaque, variegated brown. Description. LT: ♀ (n = 3): 17.3 mm (16.3–18.0). Head: red, more or less marked with green and brown (Figs. 1 G–H); frons often with brown transverse line along superior margin of disc, and oblique discal carinae green-brown; infero-lateral angles of frons and baso-lateral angles of postclypeus marked with green-brown (Fig. 1 H); antennae orange to red; pedicel large, inflated, reniform (Fig. 1 H); ratio BV/LV = 4.8; BF/LF = 1.6. Thorax: prothorax red more or less marked with green and brown, especially infero-lateral angles (Fig. 1 G); mesonotum red with brown markings on sides of disc (Fig. 1 G); ratio LP+LM/BT = 0.9; LM/LP = 2.1. Tegmina: variegated orange-brown to brown with membrane opaque, hyaline in places, with brown confluent spots; costal cell marked with pale green and 4–5 dark brown patches (Fig. 1 G); ratio LTg/BTg = 3.3. Hind wings: base largely orange limited by broad brown line extending from middle of costal margin, along basal half of sutural margin, to postero-basal angle; apex hyaline; veins brown, orange in orange zone (Fig. 1 G). Legs: red; tibiae with 1-2 green-brown rings; apices of spines of hind legs black (Fig. 1 G). Abdomen: bright red with basal tergites black (Fig. 1 G). Biology. Two out of the five specimens have been caught at light trap. Distribution (Fig. 5). Arnhem Land, Northern Territory. FULGORIDAE: DESUDABOIDES


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Discussion The discovery of the four new species greatly extends westwards and southwards the known distribution of the genus Desudaboides in Australia. The important number of new species found in recent material (all collected after 1964) compared to the single species described since 1927 indicates that other unknown species will be discovered in the future. More fieldwork is needed to obtain information on the biology of those insects which are only known to be attracted to light (13 of the 28 examined specimens). It is possible that they are nocturnal, hiding during the day under loose bark or in crevices of trunks, or in the canopy, like other species of the family (Naskrecki & Nishida, 2007).

Acknowledgments I thank the curators listed above for the loan of the material, Mr Tom Weir for transmitting the data of ANIC (Canberra, Australia), Dr Max Moulds (Kuranda, Queensland, Australia) for the information about the location of Bluewater Range, Mr Pol Limbourg and Dr Patrick Grootaert (RBINS), D. Thierry Bourgoin and Dr Adeline Soulier-Perkins (Museum National d’Histoire Naturelle, Paris, France), and Dr Murray J. Fletcher (Agricultural Scientific Collections Unit, Orange, Australia) for their support and comments. This study has also benefited from funds of the European Union for a visit to the collections of the MNHN (COLPARSYST project).

References Barbier, Y. & Rasmont, P. (2000) Carto Fauna-Flora 2.0. Guide d’utilisation. Université de Mons Hainaut, Mons, Belgique, 59 pp. Constant, J. (2004) Révision des Eurybrachidae (I). Le genre Amychodes Karsch, 1895 (Homoptera: Fulgoromorpha: Eurybrachidae). Bulletin de l’Institut royal des Sciences naturelles de Belgique, 74, 11–28. Lallemand, V. (1963) Révision des Fulgoridae (Homoptera). Deuxième partie. faunes asiatique et australienne. Mémoires de l'Institut royal des Sciences naturelles de Belgique (2e série), 75, 1-99, pl. 1–11. Metcalf, Z.P. (1947) General Catalogue of the Homoptera. Fascicle IV Fulgoroidea. Part 9 Fulgoridae. Raleigh (U.S.A.) North Carolina State College, 280 pp. Musgrave, A. (1927) Contribution to the knowledge of Australian Hemiptera. N° 1. Records of the Australian Museum, 16, 47–49, plate 4. Nagai, S. & Porion, T. (1996) Fulgoridae 2: Catalogue illustré des faunes asiatique et australienne. Sciences Nat, Compiègne: 80pp., 236 figs. Naskrecki, P. & Nishida, K. (2007) Novel trophobiotic interactions in the lantern bugs (Insecta: Auchenorrhyncha: Fulgoridae). Journal of Natural History, 41 (37–40), 2397–2402.

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