Zootaxa, Revision of the genus Eulichas Jacobson ...

Zootaxa 2192: 1–44 (2009) www.mapress.com / zootaxa/

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ZOOTAXA ISSN 1175-5334 (online edition)

Revision of the genus Eulichas Jacobson, 1913 (Coleoptera: Eulichadidae) II. E. dudgeoni species group JIŘÍ HÁJEK Department of Entomology, The National Museum, Kunratice 1, CZ-148 00 Praha 4, Czech Republic. E-mail: [email protected]

Table of contents Abstract ............................................................................................................................................................................... 2 Introduction ......................................................................................................................................................................... 2 Material and methods .......................................................................................................................................................... 2 Systematics .......................................................................................................................................................................... 3 The Eulichas dudgeoni species group ................................................................................................................................. 3 Checklist and distribution of the species ............................................................................................................................. 4 A key to species .................................................................................................................................................................. 4 The Eulichas dudgeoni species complex ............................................................................................................................ 6 Eulichas alesbezdeki Hájek, sp. nov. ........................................................................................................................... 6 Eulichas bertiae Jäch, 1995 ........................................................................................................................................ 15 Eulichas dudgeoni Jäch, 1995 .................................................................................................................................... 17 Eulichas sp. ♀♀ (bertiae / dudgeoni)........................................................................................................................ 18 Eulichas jendeki Jäch, 1995 ....................................................................................................................................... 18 Eulichas siamensis Hájek, sp. nov. ........................................................................................................................... 20 Eulichas similis Hájek, sp. nov. ................................................................................................................................. 21 Eulichas uniformis (Pic, 1911) .................................................................................................................................. 23 The Eulichas sikkimensis species complex ....................................................................................................................... 24 Eulichas baeri (Fairmaire, 1898)............................................................................................................................... 24 Eulichas gigantea (Fairmaire, 1891) ......................................................................................................................... 26 Eulichas incisicollis Pic, 1933 ................................................................................................................................... 27 Eulichas oborili Hájek, sp. nov. ............................................................................................................................... 28 Eulichas robusta Hájek, sp. nov. .............................................................................................................................. 29 Eulichas sausai Hájek, sp. nov. ................................................................................................................................ 31 Eulichas serricornis Hájek, sp. nov. ........................................................................................................................ 32 Eulichas sikkimensis (Pic, 1913) ............................................................................................................................... 34 Eulichas sundaensis Hájek, sp. nov. ........................................................................................................................ 35 Eulichas wewalkai Hájek, sp. nov. ............................................................................................................................ 36 The Eulichas fasciolata species complex ......................................................................................................................... 36 Eulichas fasciolata (Fairmaire, 1898) ....................................................................................................................... 37 Eulichas jakli Hájek, sp. nov. .................................................................................................................................... 38 Eulichas subocellata (Fairmaire, 1898) ..................................................................................................................... 38 Eulichas villosa Hájek, sp. nov. ................................................................................................................................ 40 Species not revised ............................................................................................................................................................ 41 Eulichas trapezicollis (Fairmaire, 1891) ................................................................................................................... 41 Discussion ......................................................................................................................................................................... 41 Zoogeographical considerations ........................................................................................................................................ 41 Acknowledgements ........................................................................................................................................................... 43 References ........................................................................................................................................................................ 43

Accepted by P. Johnson: 11 Jul. 2009; published: 10 Aug. 2009

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Abstract The Eulichas dudgeoni species group of the genus Eulichas Jacobson, 1913 is revised. Three informal species complexes are established. All included species are described and their diagnostic characters are illustrated. The following eleven new species are described: E. alesbezdeki Hájek, sp. nov. (Laos, Vietnam), E. jakli Hájek, sp. nov. (Indonesia: Kalimantan), E. oborili Hájek, sp. nov. (Thailand), E. robusta Hájek, sp. nov. (Malaysia), E. sausai Hájek, sp. nov. (Indonesia: Sumatra, Malaysia), E. serricornis Hájek, sp. nov. (Malaysia), E. siamensis Hájek, sp. nov. (Laos, Thailand, Vietnam), E. similis Hájek, sp. nov. (Laos, Thailand), E. sundaensis Hájek, sp. nov. (Indonesia: Sumatra, Java), E. villosa Hájek, sp. nov. (Malaysia: Sabah), and E. wewalkai Hájek, sp. nov. (Nepal). E. baeri (Fairmaire, 1898) is synonymised with E. baeri var. innotatus Pic, 1924 syn. nov. Lectotypes are designated for Eulichas baeri innotatus (Luzon, Philippines), E. incisicollis Pic, 1933 (Cameron Highlands, Malaysia), and Lichas subocellata Fairmaire, 1898 (Kinabalu, Sabah, Malaysia). Key words: Coleoptera, Eulichadidae, Eulichas, Eulichas dudgeoni species group, taxonomy, new species, Palaearctic, Oriental, identification key

Introduction The genus Eulichas Jacobson, 1913 belongs (together with the monotypic Californian genus Stenocolus LeConte, 1853) to the small elateriform family Eulichadidae. The genus comprises 31 species so far described occurring predominantly in the Oriental zoogeographical region from Nepal, northern India and southern China, through continental south-eastern Asia to the Greater Sunda Islands Sumatra, Java and Kalimantan, and the Philippines. Only a few species reach the border of the Palaearctic region in Nepal, Bhutan, northern India and China (Jäch & Hájek 2006). Larvae of Eulichas are aquatic, while adults can be collected on vegetation near streams and are attracted to light. The first modern review of Eulichas was provided by Jäch (1995), who described the new subgenus Forficulichas Jäch, 1995, established species groups, and revised taxa from China, Laos and Vietnam. Ivie & Jäch (2002) described an additional species from Vietnam. Hájek (2007, 2008) summarised known information about the genus, published a revision of the E. funebris species group, and added a new species from Laos respectively. This work represents the second part of my revision, the E. dudgeoni species group.

Material and methods In descriptions, I follow the style used in the first part of my revision (Hájek 2007), which gives only diagnostic characters usable for identification of species and variability as compared with the general description of the genus. The classification is based on males only, because the diagnostic characters predominantly apply to male genitalia. The shape of the aedeagus was studied dry, because of the membranous part of the parameres, which could be damaged when dipped. When the aedeagus is retracted, the parameres closely fit the median lobe. For studying (and drawing) the exact shape of the median lobe, the parameres were withdrawn. Thus, the angle between the parameres and the median lobe as illustrated should be regarded as artificial. The male genitalia were preserved dry on the same card as the beetles, or on separate card beneath the beetle, if it is direct-pinned. The spelling of geographical names and their coordinates were unified according to “MICROSOFT ENCARTA WORLD ATLAS 2000”. Exact label data are cited for the type material. A forward slash (/) separates different lines and a double slash (//) different labels of data. Additional remarks are found in square brackets. Holotypes of newly described species are provided with one red label with printed text: “HOLOTYPE / EULICHAS (s. str.) ♂ / “name of the species” sp. nov. / Jiří Hájek det. 2008”. Each paratype

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· Zootaxa 2192 © 2009 Magnolia Press

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is provided with a red label similar to that of the holotype, except “PARATYPE” instead of “HOLOTYPE”, the respective sex symbol and collection number. Material studied is deposited in the following institutional and private collections: BMNH CNCO DEIC EUMC HNHM IJCM IRSNB MCSN MHNG MNHN MZMB NHMB NHMW NHRS NKME NMPC RMNH SMNS SMTD SSCT VKCZ ZMAN ZMAS ZMHB ZSMC

The Natural History Museum [former British Museum], London, Great Britain (Maxwell V. L. Barclay); Canadian national collections of insects, arachnids and nematodes, Ottawa, Canada (Pat Bouchard); Deutsches Entomologisches Institut, Müncheberg, Germany (Lothar Zerche); Ehime University, Matsuyama, Japan (Hiroyuki Yoshitomi); Hungarian Natural History Museum [Magyar Természettudományi Múzeum], Budapest, Hungary (Ottó Merkl); Ivo Jeniš collection, Mladeč, Czech Republic; Institut Royal des Sciences Naturelles de Belgique, Brussels, Belgium (Jerome Constant); Museo Civico di Storia Naturale “Giacomo Doria” Genova, Italy (Roberto Poggi); Muséum d’Histoire Naturelle, Geneva, Switzerland (Giulio Cuccudoro); Muséum Nationale d’Histoire Naturelle, Paris, France (Thierry Deuve, Azadeh Taghavian); Moravské zemské muzeum, Brno, Czech Republic (Petr Baňař); Naturhistorisches Museum Basel, Switzerland (Eva Sprecher, Michel Brancucci); Naturhistorisches Museum Wien, Austria (Manfred A. Jäch); Naturhistoriska Riksmuseet, Stockholm, Sweden (Bert Viklund); Naturkundemuseum Erfurt, Germany (Matthias Hartmann); Národní muzeum, Praha, Czech Republic (Jiří Hájek); Nationaal Natuurhistorische Museum (“Naturalis”), [former Rijksmuseum van Natuurlijke Historie], Leiden, The Netherlands (Fred van Assen); Staatliches Museum für Naturkunde, Stuttgart, Germany (Wolfgang Schawaller); Staatliches Museum für Tierkunde, Dresden, Germany (Olaf Jäger); Stanislav Snäll collection, Tumba, Sweden; Vít Kabourek collection, Zlín, Czech Republic; Zoologisch Museum, Universiteit van Amsterdam, The Netherlands (Ben Brugge); Zoological Institute, Russian Academy of Science, Sankt Petersburg, Russia (Alexander G. Kirejtshuk); Museum für Naturkunde der Humboldt-Universität, Berlin, Germany (Manfred Uhlig); Zoologische Staatssammlung, München, Germany (Martin Baehr, Michael Balke).

Systematics The Eulichas dudgeoni species group The E. dudgeoni species group is characterised by a relatively broad phallobase, which is about the same length or slightly shorter than the parameres, and by the short basal parameral apophysis. In addition, the last antennomere is filiform, rectangular or slightly club shaped, never distinctly widened, its ventral part is smooth; and the last maxillary palpomere is club shaped. The group contains 21 species distributed from Nepal, northern India and southern China, through continental south-eastern Asia, where it reaches its highest diversity, to the Greater Sunda Islands and the Philippines.

REVISION OF EULICHAS II

Zootaxa 2192 © 2009 Magnolia Press ·

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Checklist and distribution of the species The E. dudgeoni species complex 1. Eulichas alesbezdeki Hájek, sp. nov. 2. Eulichas bertiae Jäch, 1995 3. Eulichas dudgeoni Jäch, 1995 4. Eulichas jendeki Jäch, 1995

5. Eulichas siamensis Hájek, sp. nov. 6. Eulichas similis Hájek, sp. nov. 7. Eulichas uniformis (Pic, 1911)

The E. sikkimensis species complex 8. Eulichas baeri (Fairmaire, 1898)

China: Yunnan; Laos: Boli Kham Xai, Hua Phan; Vietnam: “Tonkin” China: Guizhou, Fujian, Hubei, Hunan China: Guangdong, Guangxi, Fujian, Hong Kong, Hubei, Jiangxi, Shaanxi, Sichuan China: Yunnan; Laos: Hua Phan, Louangnamtha, Louangphrabang; Myanmar; Thailand: Chiang Mai, Mae Hong Son, Nan, Tak; Vietnam: “Tonkin” Laos: Louangnamtha, Louangphrabang, Oudomxai; Thailand: Chiang Mai, Nan; Vietnam: Lao Cai Laos: Louangphrabang, Oudomxai; Thailand: Chiang Mai, Mae Hong Son, Nan Bhutan?; India: Assam, Meghālaya, Sikkim, Uttar Pradesh, West Bengal; Myanmar: Kachin; Nepal

13. Eulichas sausai Hájek, sp. nov. 14. Eulichas serricornis Hájek, sp. nov. 15. Eulichas sikkimensis (Pic, 1913) 16. Eulichas sundaensis Hájek, sp. nov. 17. Eulichas wewalkai Hájek, sp. nov.

Brunei; Indonesia: Kalimantan; Malaysia: Sabah, Sarawak; Philippines: Leyte, Luzon, Mindanao, Mindoro, Negros India: Himāchal Pradesh Malaysia: Pahang, Perak Thailand: Nan Malaysia: Kelantan, Pahang, Perak, Selangor; Thailand: Nakhon Si Thammarat, Saraburi Indonesia: Sumatra; Malaysia: Pahang Malaysia: Kelantan, Pahang, Perak Bhutan?; India: Sikkim, Uttaranchal, West Bengal; Nepal Indonesia: Sumatra, Java Nepal

The E. fasciolata species complex 18. Eulichas fasciolata (Fairmaire, 1898) 19. Eulichas jakli Hájek, sp. nov. 20. Eulichas subocellata (Fairmaire, 1898) 21. Eulichas villosa Hájek, sp. nov.

Malaysia: Sabah, Sarawak Indonesia: Southern Kalimantan Malaysia: Sabah, Sarawak Malaysia: Sabah

9. Eulichas gigantea (Fairmaire, 1891) 10. Eulichas incisicollis Pic, 1933 11. Eulichas oborili Hájek, sp. nov. 12. Eulichas robusta Hájek, sp. nov.

A key to species This key is for males only. Eulichas gigantea, known only from the female holotype, is not included. It is similar to E. robusta sp. nov. in habitus. For their separation see differential diagnosis of this species. 1. 2.

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Parameres of various shapes, their apices flattened. Median lobe from slender and subparallel to lanceolate........... 2 Parameres parallel sided, with characteristic twisted apices (Figs. 81–84). Median lobe narrowly lanceolate, always exceeding parameres. Borneo ... E. fasciolata species complex................................................................................ 18 Parameres with well developed subbasal hook (cf. Fig. 65). Border area between Palaearctic and Oriental zoogeographical region (from northern India and Nepal to southern and central China, and northern part of Thailand, Laos and Vietnam) ........................................................................................................... E. dudgeoni species complex ... 3


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7.

8.

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9. 10.

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11. 12.

13. -

14.

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Subbasal parameral hook absent (cf. Fig. 72). Continental SE Asia, Sumatra, Java, Borneo, Philippines.................... ............................................................................................................................. E. sikkimensis species complex..... 9 Sides of pronotum more or less regularly rounded (Figs. 45–47) .............................................................................. 4 Sides of pronotum skewed and straight in anterior half, with obtuse angle behind the midlength, and prominent hind angles (Figs. 48–51)..................................................................................................................................................... 6 Last antennomere short, ca. 2.5–3.1 times as long as broad (Fig. 25). Ocellations on elytra distinct (Fig. 1). Northern Laos and Vietnam ..................................................................................................................... E. alesbezdeki sp. nov. Last antennomere long, at least 3.5 times as long as wide (Figs. 26–27). Ocellations on elytra rather indistinct (Figs. 2–3). China .................................................................................................................................................................. 5 Parameres slender, their inner side sinuously narrowing to subapical hook (Fig. 67) .............................. E. dudgeoni Parameres broad, their inner side almost straight (Fig. 66) ........................................................................... E. bertiae Parameres almost parallel sided, their inner side straight, with indistinct attenuation before subapical hook (Fig. 68). Median lobe slender. Last antennomere ca. 2.7–3.0 times as long as broad (Fig. 28). Sides of pronotum basally usually subparallel (Fig. 48). Widely distributed from southern China to northern Thailand, Laos and Vietnam ............. ....................................................................................................................................................................... E. jendeki Parameres with their inner side sinuously attenuating to subapical hook (Figs. 69–70). Median lobe subparallel or narrowly lanceolate. Sides of pronotum basally usually slightly skewed (Figs. 49–50) ............................................. 7 Last antennomere short, ca. 2.2–2.5 times as long as broad (Fig. 30). Elytra with distinct yellow-brownish or greyish ocellations (Fig. 6). Parameres basally very broad, median lobe lanceolate (Fig. 70). Northern Thailand and Laos ... .......................................................................................................................................................... E. similis sp. nov. Last antennomere longer, ca. 2.7–3.8 times as long as broad. Elytra usually only with indistinct grey or yellowish ocellations. Median lobe subparallel or narrowly lanceolate ...................................................................................... 8 Smaller, males ca. 17–19 mm, grey coloured species (Fig. 7). Last antennomere shorter, ca. 2.7–3.3 times as long as broad (Fig. 31). Parameres slender with relatively longer apical part, median lobe narrowly lanceolate (Fig. 71). Nepal, northern India and Bhutan ............................................................................................................. E. uniformis Larger, males ca. 23–28 mm, brown coloured species (Fig. 5). Last antennomere longer, ca. 3.1–3.8 times as long as broad (Fig. 29). Parameres broad with shorter apical part, median lobe slender, subparallel (Fig. 69). Northern Laos, Thailand and Vietnam .................................................................................................................. E. siamensis sp. nov. Sides of pronotum almost regularly rounded (cf. Fig. 55) ........................................................................................ 10 Sides of pronotum almost straight with more or less distinct obtuse angle near the midlength and prominent posterior angles (cf. Fig. 52)............................................................................................................................................... 14 Elongate subparallel species. Body colouring brown-blackish, pubescence ocellations formed as irregular dark spots (Fig. 13). Parameres parallel sided with short apical part, median lobe lanceolate (Fig. 74). Thailand: Nan province .......................................................................................................................................................... E. oborili sp. nov. Broader species, not subparallel. Body colouring brownish red to brown, pattern formed of dark transverse stripes and pale ocellations. Parameres parallel sided or narrowing to apex, their apical part longer. Median lobe subparallel or lanceolate .............................................................................................................................................................. 11 Smaller, males ca. 16–24 mm, brown coloured species. Ocellations indistinct, formed as stripes of small spots (Fig. 9). Parameres subparallel, median lobe very narrowly lanceolate (Fig. 72). Borneo............................ E. baeri partim Larger, males more than 20 mm, brownish red or brown coloured species. Pattern on elytra forming distinct ocellations. Malay Peninsula, Sumatra ............................................................................................................................... 12 Very large species, males ca. 26–33 mm. Last antennomere rectangular, ca. 3.2–3.6 times as long as broad (Fig. 35). Parameres broad, narrowing slightly to apex; median lobe broadly lanceolate (Fig. 75). Malay Peninsula ................. ......................................................................................................................................................... E. robusta sp. nov. Smaller species, males ca. 20–24 mm. Last antennomere of various shape. Parameres slender, median lobe slender, subparallel .................................................................................................................................................................. 13 Antennomeres very serrate, last antennomere filiform, ca. 4.0–5.1 times as long as broad (Fig. 37). Pattern on elytra forming distinct ocellations (Fig. 16). Parameres parallel sided (Fig. 77). Malay Peninsula ... E. serricornis sp. nov. Antennomeres less serrate, last antennomere rectangular, ca. 2.6–3.1 times as long as broad (Fig. 36). Pattern on elytra forming distinct ocellations confluent to transverse stripes (Fig. 15). Parameres broadest basally, slightly narrowing to apex (Fig. 76). Malay Peninsula, Sumatra......................................................................... E. sausai sp. nov. Sides of pronotum with distinct angle behind the middle, pronotum “incised” baso-laterally (Fig. 53). Brownish red to brown coloured species, elytral pattern with distinct ocellations (Figs. 11–12). Antenna slender, short – reach before body midlength. Last antennomere narrowly oval, ca. 2.7–3.2 times as long as wide (Fig. 33). Parameres slender, narrowing slightly to apex; median lobe slender, subparallel in basal two thirds of its length (Fig. 73). Cameron Highlands in Malay Peninsula ........................................................................................................ E. incisicollis Sides of pronotum with only minute angle near the midlength. Usually dark coloured species, with indistinct ocellations. Antenna longer, reaching body midlength. Other characteristics variable ...................................................... 15 Elongate subparallel species. Sides of pronotum anteriorly slightly rounded, angle behind midlength substituted by



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16. 17.

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18.

19. 20.

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slight sinuation, posterior angles very prominent (Fig. 59). Last antennomere filiform, ca. 4.6–6.9 times as long as broad (Fig. 39). Parameres slender, parallel sided, their apical part very short; median lobe slender, very narrowly lanceolate (Fig. 79). Sumatra, Java ............................................................................................ E. sundaensis sp. nov. Broader species, not subparallel. Sides of pronotum anteriorly skewed, angle more or less distinct. Last antennomere of variable shape. Parameres with longer apical part. Median lobe of variable shape ............................................... 16 Parameres parallel sided (Fig. 72). Philippines .................................................................................... E. baeri partim Parameres not parallel sided. Species from Bhutan, northern India and Nepal......................................................... 17 Larger species, males ca. 16–19 mm. Antenna slender, last antennomere ca. 3.8–4.3 times as long as broad (Fig. 38). Aedeagus broad, parameres attenuating to apex; median lobe narrowly lanceolate (Fig. 78). Bhutan, northern India, Nepal ...................................................................................................................................................... E. sikkimensis Smaller species, male 14 mm. Antenna shorter, last antennomere ca. 2.6 times as long as broad (Fig. 40). Aedeagus slender, parameres slightly constricted before their midlength; median lobe broadly lanceolate with distinct lateral lobes (Fig. 80). Nepal.................................................................................................................... E. wewalkai sp. nov. Smaller, slender species, male 21 mm. Antenna slender, last antennomere filiform, ca. 4.7 times as long as broad (Fig. 42). Parameres with longer apical part, apex not emarginated (Fig. 82). Indonesia: Kalimantan ........................ .............................................................................................................................................................. E. jakli sp. nov. Larger species, males usually exceed 25 mm. Antenna broader, last antennomere less than 4.5 times as long as broad. Parameres with short apical part, apex distinctly emarginated. Malaysia: Sabah, Sarawak .......................... 19 Setation on elytra forms distinct grey-whitish ocellations (Fig. 23). Last antennomere narrowly oval with apical elongation, ca. 3.4–4.5 times as long as wide (Fig. 43) ......................................................................... E. subocellata Setation on elytra yellowish, ocellations indistinct. Antenna of various shape ........................................................ 20 Larger and broader species, males 24–32 mm. Body setation comparatively sparser (Fig. 21). Antenna robust, last antennomere oval, ca. 3.2–3.6 times as long as broad (Fig. 41). Parameres parallel sided, their subapical hook very prominent (Fig. 81). Sabah, Sarawak......................................................................................................... E. fasciolata Smaller, slender and more parallel species, males 24–27 mm. Body setation dense (Fig. 24). Antenna slender, last antennomere nearly filiform, ca. 3.8–4.4 times as long as broad (Fig. 44). Parameres slightly narrowed medially, their subapical hook less prominent (Fig. 84). Sabah ....................................................................... E. villosa sp. nov.

The Eulichas dudgeoni species complex The species complex contains seven species from, more or less, the Palaearctic-Oriental transitional zone area. Their characteristics agree without exception with the definition of the group: Parameres have a distinctly developed subbasal hook. In addition, in most species parameres are distinctly attenuated before the subapical hook, and the median lobe is narrowly lanceolate and often attenuated in the apical third of its length.

Eulichas alesbezdeki Hájek, sp. nov. (Figs. 1, 25, 45, 65) Eulichas sp. (Guangxi A) cf. funebris et undulata: Jäch 1995: 370 (partim). Eulichas jendeki: Jäch 1995: 373 (partim).

Type locality. Laos, Boli Kham Xai Prov., 8 km NE of Ban Nape. Type material. 79 specimens — Holotype ♂ (NMPC), labelled: “LAOS-CE, 1–18.v.2001, / Boli Kham Xai prov., / 18°21’N 105°08’E, / BAN NAPE (8 km NE), / ~600m, Vít Kubáň leg. [printed] / Biological expedition / „Laos 2001“ / Moravian Museum Brno / Czech Republic [printed]”. Paratypes: 20♂♂ (nos. 1– 20) 6♀♀ (nos. 21–26), same label data as holotype (MZMB, NHMW, NMPC); 13♂♂ (nos. 27–39) 4♀♀ (nos. 40–43), same label data, but “L. Dembický leg.” (NHMB, NMPC); 2♂♂ (nos. 44–45), “LAOS – NE; HUA PHAN prov. / 25km SE Vieng Xai (by road); / BAN KANGPABONG env.; / 20°19’N 104°25’E; / J. Bezděk leg.; 14.–18.v.2001 [printed]” (NMPC); 1♂ (no. 46), “LAOS-C, Kamphéng Nakhon / Viangchan prov., Viang Chan / (=VIENTIANE), 17°58’N / 102°36’E, 170m, 17.v.2008, / Khampaseuth Sisoutham leg. [printed]” (NMPC); 1♂ (no. 47), “Tonkin / Than-Moi / Juni-Juli / H.Fruhstorfer [printed]” (HNHM); 6♂♂ (nos. 48–53),

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“Tonkin / Montes Mauson [Tam Dảo], / April,Mai 2-3000’ / H.Fruhstorfer [printed]” (HNHM, ZMHB); 2♂♂ (nos. 54–55), “Vietnam Tam dao / Vinh phu pr. / 3.–11.6.1985 / J. Picka lgt. [printed]” (NHMB); 1♀ (no. 56), “Vietnam / Tam dao 3.6.–11.6.1985 / Vinh phu prov. / Strnad Jan lgt. [printed]” (NMPC); 1♂ (no. 57), “Vietnam N / Tam dao 26.5.–3.6.1986 / Vinh phu prov. / Strnad Jan lgt. [printed]” (NMPC); 4♂♂ (nos. 58–61) 1♀ (no. 62), “N.VIETNAM 900m / TamDao 13.–24. / 5.1989 A. Olexa [printed]” (NMPC); 1♂ (no. 63) 3♀♀ (nos. 64–66), “VIETNAM 20.–28.VI.1990 / Vinh Phu Prov. / TAM DAO / Jan Strnad leg. [printed]” (NMPC); 1♀ (no. 67), “N. Vietnam / Tamdao / 20.6.1995 Gorochov [handwritten]” (ZMAS); 1♀ (no. 68), “N-Vietnam, Tam Dao, Vinh / Phu Prov., 21°27’18’’N; / 105°38’58’’E / 2.–6.VI.1999, 1050–1200m; / leg.: Fabrizi & Ahrens [printed]” (SMTD); 1♂ (no. 69), “Vietnam gory [Mts.] SW / Cua-rao 600 m / 24.6.196 Kabakov [printed, transcribed from Cyrillic]” (ZMAS); 1 ♂ (no. 70), “Vietnam / Lao Cai [Lào Cai, ca. 22°30’N 103°57’E] / 15.5.1960 Izokh [handwritten, transcribed from Cyrillic]” (ZMAS); 1♂ (no. 71), “Vietnam gory [Mts.] SW / Bai Tuong [Bái Thuơọơ ng] 300 m / 5.7.1962 Kabakov [printed, transcribed from Cyrillic]” (ZMAS); 1♀ (no. 72), “Vietnam gory [Mts.] / W Sa-pa [Sa Pa, ca. 22°20’N 103°50’E] 1600–2000 m / 1.6.1963 Kabakov [printed, transcribed from Cyrillic]” (ZMAS); 1♂ (no. 73), “Vietnam gory [Mts.] 50 km / NO [NE] Thai Nguyen [Thái Nguyên, ca. 21°33’N, 105°51’E] 300 m / 15.6.1963 Kabakov [printed, transcribed from Cyrillic]” (ZMAS); 1♂ (no. 74), “Vietnam gory [Mts.] 60 km / NO [NE] Thai Nguyen 300 m / 16.6.1963 Kabakov [printed, transcribed from Cyrillic]” (ZMAS); 1♀ (no. 75), “16.–17.5.1990 YEN BAI [Yên Bái] / HOANG LIEN SON Distr. / N VIETNAM / JAN HORÁK Leg. [printed]” (NMPC); 2♂♂ (nos. 76–77) 1♀ (no. 78), “China / Yunnan [printed]” (HNHM, NMPC). Additional material studied. 12 specimens — LAOS: 1♀ , Annam (HNHM). VIETNAM: 1♀ , H t Tonkin, A.N. Claire (IRSNB); 1♀, Tonkin, Rég.[ion] Hà Giang, 1916, S. Olivier (MNHN); 1♀ , Tonkin, Chapo, prov. de Laokay [Sa Pa, ca. 22°20’N 103°50’E; Lao Cai prov.] (MNHN); 1♀, Tonkin, NE of Bảo Lạc [ca. 22°57’N 105°39’E], 1897–1898, Dr. Battarel leg. (MNHN); 1♀ , Làng Son [ca. 21°51’N 106°45’E] (MNHN); 1♂, Tonkin, Than-Moi [most probably Thanh Moi, ca. 21°38’N 106°32’E], v.–vi., H. Fruhstorfer (MNHN) [paratype of E. jendeki]; 1♂, Tonkin, Chiêm Hóa, viii.–ix., H. Fruhstorfer (MNHN) [paratype of E. jendeki]; 1♂, Tonkin, Hòa Bình env. [ca. 20°50’N 105°19’E], 1902, J. Laisi leg. (MNHN) [paratype of E. jendeki]; 2♂♂, Tonkin, Rég.[ion] Hòa Bình, 1919 (MNHN) [paratypes of E. jendeki]; 1♂, Tonkin, Backan [Bảc Cạn, ca. 22°08’N 105°49’E], 1907–08, P. Lemée (NHMW) [paratype of E. jendeki]. Description. Habitus elongate, fusiform. Body colouring from brownish-red to black. Pale part of setation consists of recumbent yellowish or greyish setae forming typical ocellations on pronotum, elytra and abdominal sternites (Fig. 1). Measurements. Males: 21–26 mm (holotype 22 mm); females: 25–31 mm. Head punctation consists of sparse moderately large setigerous punctures. Antenna robust, last antennomere nearly rectangular, 2.57–3.06 times as long as wide (Fig. 25), its ventral side smooth. Pronotum trapezoidal, ca. 1.89–2.05 times as wide as long. Sides almost regularly rounded (Fig. 45). The disc with two oval shallow depressions. Punctation consists of sparse moderately coarse setigerous punctures on the disc, which become slightly coarser and denser laterally. Elytra with numerous longitudinal rows of moderately large punctures, and very fine interstitial setigerous punctures. Ventral part with fine punctures, which are sparse medially and become larger and denser laterally. Last abdominal ventrite laterally regularly rounded to apex. Male. Aedeagus with phallobase shorter than parameres. Parameres simple, relatively slender, their subbasal hook small, but the subapical hook is well developed and prominent. Median lobe narrowly lanceolate (Fig. 65). Female. Similar to male in habitus, but larger. Antenna shorter, and more slender. Differential diagnosis. The new species is similar to E. jendeki in habitus, while its aedeagus is rather close to that of E. dudgeoni. Eulichas alesbezdeki sp. nov. differs from E. jendeki in the almost regularly rounded sides of pronotum and the slender and slightly attenuating parameres of aedeagus, and from E. dudgeoni in shorter last antennomere and distinct body ocellations. REVISION OF EULICHAS II


7

FIGURES 1–12. Habitus of Eulichas. 1—E. alesbezdeki sp. nov. (holotype); 2—E. bertiae (paratype, Suisapa); 3—E. dudgeoni (paratype, Hong Kong); 4—E. jendeki (Sa Pa); 5—E. siamensis sp. nov. (paratype, Ban Bo Klua); 6—E. similis sp. nov. (paratype, Soppong); 7—E. uniformis (Dolakha); 8—E. baeri (Luzon); 9—E. baeri (Crocker Range); 10—E. gigantea (holotype); 11—E. incisicollis ♂ (Tanah Rata); 12—E. incisicollis ♀ (holotype).

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HÁJEK

FIGURES 13–24. Habitus of Eulichas. 13—E. oborili sp. nov. (holotype); 14—E. robusta sp. nov. (holotype); 15—E. sausai sp. nov. (holotype); 16—E. serricornis (paratype, Felda Lasah); 17—E. sikkimensis (Danda Pakhar); 18—E. sundaensis sp. nov. (holotype); 19—E. sundaensis sp. nov. ♀ (Java); 20—E. wewalkai sp. nov. (holotype); 21—E. fasciolata (Crocker Range); 22—E. jakli sp. nov. (holotype); 23—E. subocellata (Tawau); 24—E. villosa sp. nov. (paratype, Gunung Emas).



9

FIGURES 25–44. Male antennomeres IX–XI of Eulichas. 25—E. alesbezdeki sp. nov.; 26—E. bertiae; 27—E. dudgeoni; 28—E. jendeki; 29—E. siamensis sp. nov.; 30—E. similis sp. nov.; 31—E. uniformis; 32—E. baeri; 33—E. incisicollis; 34—E. oborili sp. nov.; 35—E. robusta sp. nov.; 36—E. sausai sp. nov.; 37—E. serricornis sp. nov.; 38— E. sikkimensis; 39—E. sundaensis sp. nov.; 40—E. wewalkai sp. nov.; 41—E. fasciolata; 42—E. jakli sp. nov.; 43—E. subocellata; 44—E. villosa sp. nov. Scale bar 1 mm.

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HÁJEK

FIGURES 45–54. Pronotum of Eulichas. 45—E. alesbezdeki sp. nov.; 46—E. bertiae; 47—E. dudgeoni; 48—E. jendeki; 49—E. siamensis sp. nov.; 50—E. similis sp. nov.; 51—E. uniformis; 52—E. baeri (Luzon); 53—E. incisicollis; 54—E. oborili sp. nov. Not in Scale.



11

FIGURES 55–64. Pronotum of Eulichas. 55—E. robusta sp. nov.; 56—E. sausai sp. nov.; 57—E. serricornis sp. nov.; 58—E. sikkimensis; 59—E. sundaensis sp. nov.; 60—E. wewalkai sp. nov.; 61—E. fasciolata; 62—E. jakli sp. nov.; 63—E. subocellata; 64—E. villosa sp. nov. Not in Scale.

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HÁJEK

FIGURES 65–71. Aedeagus of Eulichas in dorsal view. 65—E. alesbezdeki sp. nov.; 66—E. bertiae; 67—E. dudgeoni; 68—E. jendeki; 69—E. siamensis sp. nov.; 70—E. similis sp. nov.; 71—E. uniformis. Scale bar 1 mm.



13

FIGURES 72–77. Aedeagus of Eulichas in dorsal view. 72—E. baeri; 73—E. incisicollis; 74—E. oborili sp. nov.; 75— E. robusta sp. nov.; 76—E. sausai sp. nov.; 77—E. serricornis sp. nov. Scale bar 1 mm.

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HÁJEK

FIGURES 78–80. Aedeagus of Eulichas in dorsal view. 78—E. sikkimensis; 79—E. sundaensis sp. nov.; 80—E. wewalkai sp. nov. Scale bar 1 mm.

Collection circumstances. Collected at light. Distribution. Known from several localities in north-eastern Laos, and “Tonkin”, northern Vietnam. Etymology. The new species is dedicated to my friend Aleš Bezděk (České Budějovice, Czech Republic), specialist on Scarabaeoidea, with my special thanks for giving me the first Eulichas specimen and stimulating my interest in this family.

Eulichas bertiae Jäch, 1995 (Figs. 2, 26, 46, 66) Eulichas bertiae Jäch, 1995: 374 (original description, Rég. de Pin-Fa); Jäch & Hájek 2006: 455 (catalogue). Type locality. “Rég. de Pin-Fa (= Pingba, Anshun Prefecture, ca. 50 km SW Guiyang City, Guizhou, southern China).” Type material. 5 specimens — Holotype ♂ (MNHN): “MUSEUM PARIS / KOUY-TSCHÉOU / RÉG. DE PIN-FA / PÈRE CAVALERIE 1909 [printed] // Muséum Paris / Coll. M. Pic [printed] // HOLOTYPE / Eulichas / bertiae sp.n. / des. M. Jäch 1994 [red label, printed]”. Paratypes: 1♂ , same data as holotype (NHMW); 2♂♂ 1♀, “Suisapa, 1000 M. / Lichuan Distr. / W. Hupeh, China / VIII-30-48 // Gressitt & / Djou Collrs. [printed]” (NHMW). Additional material studied. 3 specimens — CHINA: 1 ♂ , Fujian prov., Kuatun [= Guadun], 7.ix.[19]46, Tschung-Sen leg. (MNHN); 1♂, Hubei prov., Lao-ho-k’ou Pr. [Laohekou distr.], Tsch’ia-yuenkow, G. Hauser leg. (ZMHB); 1♂, Hunan prov., Wulingyuan, N Dayong, Zangjiajie, 450 m, 30.x.1993, H. Schillhammer leg. (NHMW).



15

FIGURES 81–84. Aedeagus of Eulichas in dorsal view. 81—E. fasciolata; 82—E. jakli sp. nov.; 83—E. subocellata; 84—E. villosa sp. nov. Scale bar 1 mm.

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HÁJEK

Description. Habitus elongate, fusiform. Body colouring black, rarely dark brown. Pale part of setation consists of recumbent yellowish or greyish setae forming rather indistinct ocellations on pronotum, elytra and abdominal sternites (Fig. 2). Measurements. Males: 20–24 mm; female: 25 mm. Head punctation consists of sparse moderately large setigerous punctures. Antenna long, slender, last antennomere almost rectangular, with apical elongation of variable length, ca. 3.60–4.64 times as long as wide (Fig. 26), its ventral side smooth. Pronotum trapezoidal, ca. 1.85–1.95 times as wide as long. Sides of rather variable shape; in some specimens almost regularly rounded (Fig. 46). On the other hand some specimens have sides skewed and straight in anterior half, subparallel in basal half, with an indistinct obtuse angle present near the middle. Disc with two oval shallow depressions. Punctation consists of sparse moderately coarse setigerous punctures on the disc, which become slightly coarser and denser laterally. Elytra with numerous longitudinal rows of moderately large punctures, and fine interstitial setigerous punctures. Ventral part with fine punctures, which are sparse medially and become larger and denser laterally. Last abdominal ventrite laterally regularly rounded to apex. Male. Aedeagus with phallobase slightly shorter than parameres. Parameres simple, relatively broad. Subbasal parameral hook small, but the subapical hook is well developed and prominent. Median lobe narrowly lanceolate (Fig. 66). Female. Similar to male in habitus, but larger. Antenna shorter, and more slender. Distribution. Known from several localities in southern and south-eastern China (Fujian, Guizhou, Hubei, Hunan). Eulichas dudgeoni Jäch, 1995 (Figs. 3, 27, 47, 67) Eulichas dudgeoni Jäch, 1995: 372 (original description, Hongkong: Tai Po Kau forest); Jäch & Hájek 2006: 455 (catalogue).

Type locality. “Stream in Tai Po Kau forest, New Territories, Hong Kong”. Type material. 73 specimens — Holotype ♂ (NHMW): “Hong Kong / Tai Po Kau / leg. Dudgeon [handwritten] / HOLOTYPUS / Eulichas / dudgeoni sp. n. / des. M. Jäch 1993 [red label, printed]”. Paratypes: 46♂♂ 2♀♀, same data, but printed (NHMW); 9♂♂ 4♀♀, “HONGKONG / leg.Dudgeon [printed] (NHMW); 2♂♂, “HONGKONG, Tai Po Kau / 10.8.1983, light / leg. Dudgeon [printed]” (NHMW); 1♂, “Coll.Nonfried / China [white label with black frame, printed] // Collect. / Plason [printed]” (NHMW); 5♂♂, “Coll.R.I.Sc.N.B. / Chine: Kwang - Si / Nan - ning / ex coll. Le Moult [printed]” (IRSNB, NHMW); 2♂♂, “Suisapa, 1000 M. / Lichuan Distr. / W. Hupeh, China / VIII-30-48 // Gressitt & / Djou Collrs. [printed]” (NHMW). Additional material studied. 23 specimens — CHINA: 1♂ 1♀, Sichuan Prov., Yannsien [= Ya’an, ca. 29°59’N 102°59’E], coll. Le Moult (IRSNB); 1♂ , Guangxi A.R., Jaochan, G. Sin leg. (ZMHB); 4 ♂♂ , Guangxi A.R., Region Nanning, 1931 (MNHN); 1♂, Guangxi A.R., Yaosan [= Dayaoshan], 17.vii.[19]34, H.G. Tao leg. (MNHN); 1♂, Guangxi A.R., Ku-ling [Guling, ca. 23°38’N, 108°17’E], 25.ii.[19] 35, O. Piel leg. (MNHN); 1 ♂ , Guangdong Prov., Lung-zo-Aam bei Kanton [= near Guangzhou] (ZMHB); 1 ♀ , Guangdong Prov., Tsha-jiu-san [= Tshayuen Shan, ca. 24°53’N 113°50’E], vii.-ix.1910, S.V. Mell leg. (ZMHB); 1♂, Guangdong Prov., Canton [= Guangzhou], Fung-wan, 1.v.1911, S.V. Mell leg. (ZMHB); 2♂♂ 1♀, Hong Kong (MNHN, NMPC); 1♂ Hongkong, Pak Sha O, 100 m, 4.viii.1989, W.J. Tennent (BMNH); 1♂ 1♀, Jiangxi prov., Kiukiang [Jiujiang, ca. 29°43’N 115°58’E]; vii.1887, A.E. Pratt leg. (MNHN); 1♀, Fujian prov., Shaowu env., 23.-27.vi.1991 (NHMW); 1♂ , Zhejiang prov., Wenchow [Wenzhou, ca. 28°00’N 120°38’E] (MNHN); 1 ♀ , Zheijang prov., Lin-An, Xi-Tianmu Shan Mt., 300-400 m, 17.vii.2000, N. Ohbayashi & L.-Z. Li leg. (EUMC); 1♂ 1♀, Shanghai (MNHN). REVISION OF EULICHAS II


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Description. Habitus elongate, fusiform. Body colouring from brownish-red to brown. Pale part of setation consists of recumbent yellowish or greyish setae uniformly covering body surface, or forming indistinct ocellations on elytra (Fig. 3). Measurements. Males: 20–25 mm; females: 25–31 mm. Head punctation consists of irregularly distributed sparse moderately large setigerous punctures. Antenna long, slender, last antennomere slightly club shaped with apical elongation of variable length, ca. 3.55–4.53 times as long as wide (Fig. 27), its ventral side smooth. Pronotum trapezoidal, ca. 1.89–2.00 times as wide as long. Sides almost regularly rounded, rarely in some specimens with very indistinct obtuse angle near the middle (Fig. 47). Pronotum with two rounded shallow depressions on the disc. Punctation consists of sparse moderately coarse setigerous punctures on the disc, which become slightly coarser and denser laterally. Elytra with numerous longitudinal rows of moderately large punctures, and very fine interstitial setigerous punctures. Ventral part with fine punctures, which are sparse medially and become larger and denser laterally. Last abdominal ventrite laterally almost regularly rounded to apex. Male. Aedeagus with phallobase slightly shorter than parameres. Parameres simple, slender, attenuating regularly to the subapical hook. Parameral subbasal hook small, but the subapical hook is well developed, slightly prominent. Median lobe slender, subparallel (Fig. 67). Female. Similar to male in habitus, but larger. Antenna shorter, and more slender. Collection circumstances. Imagos collected mostly at light (cf. Jäch 1995). Distribution. A species occurring in central and southern China (Fujian, Guangdong, Guangxi Hong Kong, Hubei, Jiangxi, Sichuan, Shaanxi). Imprecisely labelled specimens are known also from “North China”. Eulichas sp. ♀♀ (bertiae / dudgeoni) Material studied. 2 specimens — CHINA: 1♀, Fujian prov., Kuatun [= Guadun], iv.-vi.1946, Tshung Sen leg. (SMNS); 1♀, Gansu prov., Bikou, 32°32’N 104°38’E, 3.-7.vii.1997, E. Kučera leg. (NMPC). Remarks. E. bertiae and E. dudgeoni represent extremely similar species in general appearance. Whereas I was not able to find any diagnostic characters on females of these two species, I am not able to resolve specific placement of specimens listed above.

Eulichas jendeki Jäch, 1995 (Figs. 4, 28, 48, 68) Eulichas jendeki Jäch, 1995: 373 (original description, Vietnam: Tam Dao National Park); Jäch & Hájek 2006: 455 (catalogue).

Type locality. “Tam Dao National Park, 75 km NW Hanoi, North Vietnam (collected at light)”. Type material. 16 specimens — Holotype ♂ (NHMW): “N-Vietnam 15.V.-16.VI. / TAMDAO N.P. / 75 km NW Hanoi / leg. E. Jendek 1991 [printed] / HOLOTYPUS / Eulichas / jendeki sp. n. / des. M. Jäch 1993 [red label, printed]”. Paratypes: 1♂ , same data as holotype (NHMW); 13 ♂♂, “N-Vietnam 25.V.-10.VI. / SAPA (Lao Cai) / 22°20’N 103°50’E / leg. E. Jendek 1991 [printed]” (IRSNB, NHMW); 1♂, “DJOUKOULA / YUNNAN / Coll.de Touzalin [black frame, printed] // 2 [handwritten] // Muséum Paris / Coll. M. Pic [printed] // L. uniformis / (defloré) [handwritten]” (MNHN). Additional material studied. 177 specimens — LAOS: 1♂ 1♀, Luang Prabang à Theng, 1888, A. Pavie leg. (MNHN) [paralectotypes of E. phoca]; 22♂♂ 9♀♀, Louangnamtha prov., 20 km NW Louang Namtha,

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HÁJEK

21°09,2’N 101°18,7’E, 800–1000 m, 5.–11.v.1997, E. Jendek & O. Šauša leg. (NHMW, NMPC); 5♂♂ 2♀♀, Louangnamtha prov., Namtha – Muang Sing, 21°09’N 101°19’E, 900–1200 m, 5.–31.v.1997, V. Kubáň leg. (NHMW, NMPC); 1♂, Hua Phan prov., 25 km SE Vieng Xai (by road), Ban Kangpabong env., 20°19’N 104°25’E, 14.–18.v.2001, D. Hauck leg. (NMPC); 1♂ , Hua Phan prov., Phu Loei Nat. Park, Ban Sakok, 20°10’N 103°12’E, 23.–26.v.2001, J. Bezděk leg. (NMPC); 1♂, same label data, but D. Hauck leg. (NMPC); 1♂, Hua Phan prov., Phu Phan Mt. Env., Ban Saluei, 20°13’N 103°59’E, 1300–2000 m, 16.–18.v.2004, F. & L. Kantner leg. (NMPC); 1♂, same label data, but 20°12’-13.5’N 103°59.5’-104°01’E, 1340–1870 m, 15.iv.– 15.v.2008, Lao collectors leg. (NMPC). THAILAND: 1♂, Tak prov., Umphang distr., Thung Yai Wildlife Sanctuary, Song Bae stream, 15°28’N 98°48’E, 300 m, light trap, 18.–27.iv.1988, M.J.D. Brendell leg. (BMNH); 1♂ , Tak prov., Région de Umphang, v.1991 (VKCZ); 3 ♂♂ , Chaing Rai prov., Wiangpapao, 26.vii.1990 (EUMC); 3♂♂ 5♀♀, Mae Hong Son prov., Ban Huai Po, 1600–2000 m, 9.–16.v.1991, J. Horák leg. (NMPC); 4♂♂, [Mae Hong Son prov.], Soppong, 19°22’N 98°18’E, 750 m, 13.v.1993, L. Bocák leg. (SMNS); 2♂♂ , same data, but 1.–9.v.2000, P. Viktora leg. (NMPC); 1♂, [Mae Hong Son prov.], Pai, 15.– 16.v.1999, R. Grimm leg. (SMNS); 1♂, Chiang Mai, viii.1996, Wong Tet Fatt leg. (HNHM); 1♂, Nan prov., Bo Klua, 19°08’N 101°10’E, 700 m, 22.–26.iv.1999, D. Hauck leg. (NMPC); 1♂, same label data, but M. Říha leg. (NMPC); 1♂, Nan prov., Ban Bo Klua env., 13.–26.v.2002, P. Průdek & M. Obořil leg. (NMPC); 1♂, Nan prov., Khun Nan Nat. Park, 23.iv.2004, P. Viktora leg. (NMPC). VIETNAM: 1♂, gory u [mountains near] Sa Pa, 1200 m, 23.v.1963, Kabakov leg. (ZMAS); 20♂♂ 8♀♀ , Lao Cai prov., Sa Pa [ca. 22°20’N 103°50’E], 11.–15.v.1990, P. Pacholátko leg. (NHMB); 9 ♂♂ 6♀♀ , same label data, but V. Kubáň leg. (NHMB, NMPC); 1♂, same label data, but J. Picka leg. (NHMB); 1♀, same label data, but 11.–18.vi.1990, A. Olexa leg. (NMPC); 6♀♀, same label data, but 25.v.–10.vi.1991, E. Jendek leg. (NHMW); 4♂♂ 2♀♀, same label data, but 1500 m, 11.–19.vi.1991, J. Strnad leg. (NHMB); 1♂, same label data, but S. Brantlová leg. (NHMB); 1♂, same data, but Hoang Lien Son Nat. Res., 1250–1300 m, 15.–20.vi.1998, A. Napolov leg. (ZMAS); 28♂♂ 9♀♀, Vinh Phuc prov., Tam Dảo [ca. 21°27’N 105°39’E], 6.–21.v.1990, P. Pacholátko leg. (NHMB, NMPC); 3♂♂ 1♀, same label data, but L. Dembický leg. (NHMB, NMPC); 1♂, same label data, but V. Kubáň leg. (NHMB); 1♂ 1♀, same label data, but 6.–25.v.1990, O. Šauša leg. (NHMB). The following specimens were identified with doubt: MYANMAR: 1♂, Hte Birmanie, Etat de Momeit [= Möng Mit, Shan state], 600 m, 1890, Doherty leg. (MNHN); 1♂, Dawna, 20.iv.1991 (SMNS); 5♂♂ 6♀♀, same label data, but 1.iv.–2.v.1992 (EUMC, NMPC). Doubtful localisation: 1♂ 1♀, Malaysia, Fraser’s Hill, 18.iv.1990, Y. Hori leg. (EUMC). Description. Habitus elongate, fusiform. Body colouring from brownish-red to brown. Pale part of setation consists of recumbent yellow greyish, or whitish setae forming indistinct ocellations on pronotum, and characteristic ocellations on elytra (Fig. 4). Measurements. Males: 19–25 mm; females: 24–28 mm. Head punctation consists of irregularly distributed sparse moderately large setigerous punctures. Antenna robust, last antennomere nearly rectangular, 2.71–3.00 times as long as wide (Fig. 28), its ventral side smooth. Pronotum trapezoidal, ca. 1.82–2.00 times as wide as long. Sides skewed and straight in anterior half, then with obtuse angle, and sides usually subparallel in basal half; hind angles very slightly prominent (Fig. 48). Pronotum with two oval shallow depressions on the disc, and indistinct depressions also medially along anterior and posterior margin. Punctation consists of rather irregularly distributed sparse moderately coarse setigerous punctures. Elytra with numerous longitudinal rows of moderately large punctures, and very fine interstitial setigerous punctures. Ventral part with fine punctures, which are sparse medially and become larger and denser laterally. Last abdominal ventrite laterally almost regularly rounded to apex. Male. Aedeagus with phallobase of the same length as parameres. Parameres simple, moderately broad, almost parallel sided, usually only slightly attenuating before subapical hook. Parameral subbasal hook small, but the subapical hook is well developed, slightly prominent. Median lobe slender, subparallel (Fig. 69). Female. Similar to male in habitus, but larger. Antenna shorter, and more slender. REVISION OF EULICHAS II


19

Remarks. The specimens from Myanmar agree with typical specimens in habitus, but differ slightly in the shape of the parameres. However, because I have not been able to find constant differences, I leave them within E. jendeki. Collection circumstances. Collected at light. Distribution. Widely distributed Oriental species, known from northern parts of Myanmar, Thailand, Laos and Vietnam, and southern China (Yunnan province).

Eulichas siamensis Hájek, sp. nov. (Figs. 5, 29, 49, 69) Eulichas dudgeoni: Jäch 1995: 373 (partim, Vietnam).

Type locality. Thailand, Nan Prov., Doi Phu Kha Nat. Park, ca. 19°13’N, 101°07’E. Type material. 96 specimens — Holotype ♂ (NMPC), labelled: “THAI, N, Nan prov., Doi / Phu Kha N.P., Headq., / 19°13’N, 101°07’E, 22– / 26.iv.1999, M. Říha leg. [printed]”. Paratypes: 2♂♂ (nos. 1–2), same label data, but “D. Hauck leg.” (NMPC); 1♂ (no. 3), “THAILAND / PROV. NAN, BO KHUA / 19. 4 – 7. 5. 2004 / MORAVEC PETR [printed]” (NMPC); 1♂ (no. 4), “THAILAND, Nan Prov. / Khun Nan NP / P. Viktora leg., 23.iv.2004 [printed]” (NMPC); 2♂♂ (nos. 5–6) 1♀ (no. 7), “N. Thailand, / Chiang Mai, / VIII. 1996, leg. Wong Tet Fatt [printed]” (HNHM); 1♂ (no. 8), “THAI-N, 1.–19.v.1998 / Chiang Mai prov., / BAN SAN PAKTIA, 1400 m, / Ivo Martinů leg. [printed]” (NMPC); 10♂♂ (nos. 9–18), “LAOS north, 5–11.V.1997 / 20 km NW Lounag Namtha, / N 21°09.2, E 101°18.7; / alt. 900± 100 m, / E. Jendek & O. Šauša leg. [printed]” (NMPC); 10♂♂ (nos. 19–28) 2♀♀ (nos. 29–30), “LAOS, Louangnamtha pr. / 21°09’N 101°19’E, / Namtha →Muang Sing, / 5 – 31. V. 1997, 900- / Vít Kubáň leg. -1200 m [printed]” (NHMW, NMPC); 1♂ (no. 31), “LAOS, Louangnamtha pr., / 21°09’N 101°19’E, / Namtha →Muang Sing, / 5–31.v.1997, 900–1200m, / Vít Kubá ň leg. [printed]” (NMPC); 1 ♂ (no. 32), “LaosPDR: Luang Namtha Prov. / Nam Ha NBCA: Lakkhammai Village / Nam Leung stream, 749mao / 47Q 074462, UTM 2339873 / 30 IV 2005, Light trap, loc 29 / N Jönsson, T Malm & B Viklund leg. [printed]” (NHRS); 2♂♂ (nos. 33–34), “LAOS: Louang Phrabang prov. / 20°33.4’N 102°14’E / 5km W Ban Song Cha / ca. 1200m, 24.4.–16.5.1999 / leg. C. Holzschuh [printed]” (NHMW); 11♂♂ (nos. 35–45), “LAOS-N, 24.iv.–16.v.1999, / Louang Phrabang prov., / 20°33-4’N 102°14’E, / Ban Song Cha (5km W), / ± 1200m, Vít Kubáň leg. // Vít Kubáň expedition / „Laos 1999“ / Moravian Museum Brno / Czech Republic [printed]” (MZMB, NMPC); 48♂♂ (nos. 46–93) 2♀♀ (nos. 94–95), “LAOS-N (Oudomxai) / 1–9.V.2002, ~1100m, / 20°45’N 102°09’E, / OUDOM XAI (17km / NEE), Vít Kubáň leg. // Entomological expedition / „Laos 2002“ / Moravian Museum / Brno, Czech Republic [printed]” (BMNH, MZMB, NMPC). Additional material studied. 37 specimens — THAILAND: 1♂, no additional data (SMNS); 4♂♂ 1♀, Chantaburi prov., Kao Soi Dao, iv. 1984, S. Steinke leg. (MCSN, NMPC); 19♂♂ 4♀♀, Chiang Mai prov. Doi Pui, v.1985, S. Steinke leg. (MCSN, NMPC); 1♂, same label data, but 3.vi.1985 (SMNS); 1♂, Doi Ithanon, 20.v.1988, G. Minet leg. (NHMB). VIETNAM: 1♂, Hoang Lien Son prov., Sa Pa, 1550 m, 11.–19.vi.1990, J. Strnad leg. (NMPC); 2♂♂, Sa Pa (Lao Cai), 27°20’N 103°50’E, 25.v.–.10.vi.1991, E. Jendek leg. (NHMW); 1♂ , Fan Si Pan Mts., Sa Pa, 1600 m, 20.iv.1995 (BMNH); 1♂ 1♀, Mont Bavi, v.1935, S. Masseyeff leg. (BMNH). Description. Habitus elongate, fusiform. Body colouring from brownish-red to brown-blackish. Pale part of setation consists of recumbent yellowish setae forming indistinct ocellations on pronotum, and characteristic ocellations on elytra (Fig. 5). Measurements. Males: 23–28 mm (holotype 24 mm); females: 29–35 mm. Head punctation consists of irregularly distributed sparse moderately large setigerous punctures. Antenna slender, last antennomere nearly rectangular, with apical elongation of variable length, 3.07–3.79 times as long as wide (Fig. 29), its ventral side smooth.

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Pronotum trapezoidal, ca. 1.87–2.02 times as wide as long. Sides skewed and straight in anterior half, then with obtuse angle, and prominent hind angles (Fig. 49). Pronotum with two oval shallow depressions on the disc, and indistinct depressions also medially along anterior and posterior margin. Punctation consists of moderately coarse setigerous punctures, sparse on the disc, and densely distributed laterally. Elytra with numerous longitudinal rows of moderately large punctures, and very fine interstitial setigerous punctures. Ventral part with fine punctures, which are sparse medially and become larger and denser laterally. Last abdominal ventrite laterally usually with more or less distinct sinuation before apex. Male. Aedeagus with phallobase shorter than parameres. Parameres simple, moderately broad, their inner side attenuating sinuously to subapical hook. Parameral subbasal hook small, but the subapical hook is well developed, very slightly prominent. Median lobe slender, subparallel (Fig. 69). Female. Similar to male in habitus, but larger. Antenna shorter, and more slender. Remarks. Some specimens from northern Thailand (Chiang Mai) display slight variability in aedeagal characters – they have parameres with ventral side almost straight, and median lobe slender than in typical specimens. Although I believe, that it is only intraspecific variability, I do not designate those specimens as paratypes. On the other hand, specimens from Sa Pa (Vietnam) were already introduced in Jäch (1995) as E. dudgeoni. Although they agree well with E. siamensis sp. nov. in habitus, shape of pronotum and last antennomere, they have parameres slightly slender than typical specimens, with their apical part more elongate, and subapical parameral hook more prominent – in these characters, they indeed resemble E. dudgeoni. They may represent a separate taxon, however, as only four specimens are known, I leave them within E. siamensis sp. nov., until more material is available. Differential diagnosis. Eulichas siamensis sp. nov. is similar to E. jendeki and E. similis sp. nov. in habitus. It differs from both of them in the longer last antennomere and more prominent hind angles of the pronotum. In addition, it differs from E. jendeki in the sinuous ventral parameral side, and from E. similis sp. nov. in the slender and subparallel median lobe of the aedeagus. Collection circumstances. Collected at light. Distribution. A species ranging in the upland area of northern Thailand, Laos and Vietnam. Etymology. The name is derived from “Siam”, an ancient kingdom spreading in the territory of recent Thailand.

Eulichas similis Hájek, sp. nov. (Figs. 6, 30, 50, 70) Type locality. Thailand, Mae Hong Son Prov., Soppong. Type material. 105 specimens — Holotype ♂ (NMPC), labelled: “Thailand bor. / Mae Hong Son prov. / Soppong / 1.–9.5.2000 / P. Viktora lgt. [printed]”. Paratypes: 1♀ (no. 1), same label data as holotype (NMPC); 15♂♂ (nos. 2–16) 9♀♀ (nos. 17–25), “THAI 13.5. 1993 / 19 29N 98 18E / SOPPONG 750 m / L. Bocák lgt. [printed]” (SMNS); 5♂♂ (nos. 26–30) 1♀ (no. 31), same label data, but “Vít Kubáň leg.” (NHMB, SMNS); 1♂ (no. 32), “THAI, N, Mae Hong Son / prov., Soppong env., 600m / 19°27’N, 98°20’ E, 28.v.– / 2.vi. 1999,D.Hauck leg. [printed]” (NMPC); 2♂♂ (nos. 33–34) 1♀ (no. 35), “Thailand NW / Mae Hong Son prov. / Soppong vill. env. / 29.4. – 17.5. 2007 / P. Viktora lgt. [printed]” (NMPC); 2♀♀ (nos. 36–37), “THAILAND bor. occ. / PAI / SOPPONG / 28. 5. – 5. 6. 1997 / Lgt. M. Snížek [printed]” (NMPC); 1♂ (no. 38) 1♀ (no. 39), “NW Thailand 23.–31.V. / Mae Hong Son 1992 / Ban Si Lang 1200m / J. Horák leg. [printed]” (NHMW); 1♂ (no. 40) 1♀ (no. 41), “NW THAILAND, 1200 m, / Mae Hong Son pr., / BAN SI LANG, 20.–22. v. / Sv. Bílý leg., 1996 [printed]” (NMPC); 1♂ (no. 42), “NW THAILAND, 30.4.- / Mae Hong Son, 4.5.1991 / Ban Huai Po, 1600–2000m / J. Horák leg. [printed]” (NMPC); 2♂♂ (nos. 43–44), “Chaing Mai / Thailand / V. 1989 [printed]” (EUMC); 1♂ (no. 45), “Wiang Papao / Thailand / VII–1992 [printed]” (EUMC); 4♂♂ (nos. 46– 49), “THAI-N,25.iv.–7.v.1996 / Chiang Mai prov., 19°19’N 98°50’E, / (Ban) SAN PAKTIA, / L.Hovorka REVISION OF EULICHAS II


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leg.,~1400m [printed]” (NMPC); 9♂♂ (nos. 50–58) 9♀♀ (nos. 59–67), “N. Thailand, / Chiang Mai, VIII. 1996, leg. / Wong Tet Fatt [printed]” (HNHM); 3♂♂ (nos. 68–70), “Thailand NE / Nan prov. - Bo Klua / 1. – 11. 5. 2001 / P. Viktora lgt. [printed]” (NMPC); 2♂♂ (nos. 71–72), “THAILAND 13.–26.V.2002 / NAN prov. / BAN BO KLUA env. / P. Průdek & M. Obořil leg. [printed]” (NMPC); 2♂♂ (nos. 73–74) 3♀♀ (nos. 75–77), “THAILAND / PROV. NAN, BO KHUA / 19. 4. – 7. 5. 2004 / MORAVEC PETR [printed]” (BMNH, NMPC); 3♂♂ (nos. 78–80) 5♀♀ (nos. 81–85), “THAILAND, Nan prov. / Khun Nan NP / P. Viktora leg., 23.iv.2004 [printed]” (NMPC); 4♂♂ (nos. 86–89) 1♀ (no. 90), “THAILAND NE / Nan province / Khun Nan National Park / 27.4.2004 / P. Viktora lgt. [printed]” (NMPC); 1♀ (no. 91), “THAILAND / CHIANG DAO / 27.5.–2.6.2002 / leg. B. Makovský [printed]” (NMPC); 4♂♂ (nos. 92–95) 5♀♀ (nos. 96–100), “LAOS-N, 24.iv.–16.v.1999, / Louang Phrabang prov., / 20°33-4’N 102°14’E, / Ban Song Cha (5 km W), / ± 1200m, Vít Kubáň leg. // Vít Kubáň expedition / „Laos 1999“ / Moravian Museum Brno / Czech Republic [printed]” (MZMB, NMPC); 1♀ (no. 101), “LAOS: Louang Phrabang prov. / 20°33.4’N 102°14’E / 5km W Ban Song Cha / ca. 1200m, 24.4.–16.5.1999 / leg. C. Holzschuh [printed]” (NHMW); 2♂♂ (nos. 102–103), “LAOS-N (Oudomxai) / 1–9.V.2002, ~1100m, / 20°45’N 102°09’E, / OUDOM XAI (17km / NEE), Vít Kubáň leg. // Entomological expedition / „Laos 2002“ / Moravian Museum / Brno, Czech Republic [printed]” (NMPC); 1♀ (no. 104), “LAOS / PAK BENG / 9. – 13. 5. 2005 / B.MAKOVSKY lgt. [printed]” (NMPC). Additional material studied. Doubtful localisation: 1♂ 4♀♀, Malaysia, Pahang, Cameron Highlands, Tanah Rata, v.1996, Wong Tet Fatt (HMNH). Description. Habitus elongate, fusiform. Body colouring from brownish-red to brown. Pale part of setation consists of recumbent yellowish or greyish setae forming indistinct ocellations on pronotum, and characteristic ocellations on elytra (Fig. 6). Measurements. Males: 21–26 mm (holotype 23 mm); females: 25–31 mm. Head punctation consists of irregularly distributed sparse moderately large setigerous punctures. Antenna robust, last antennomere nearly rectangular, 2.17–2.44 times as long as wide (Fig. 30), its ventral side smooth. Pronotum trapezoidal, ca. 1.87–2.00 times as wide as long. Sides skewed and straight in anterior half, then with obtuse angle, and prominent hind angles (Fig. 50). Pronotum with two oval shallow depressions on the disc, and indistinct depressions also medially along anterior and posterior margin. Punctation consists of rather irregularly distributed sparse moderately coarse setigerous punctures. Elytra with numerous longitudinal rows of moderately large punctures, and very fine interstitial setigerous punctures. Ventral part with fine punctures, which are sparse medially and become larger and denser laterally. Last abdominal ventrite laterally regularly rounded to apex. Male. Aedeagus with phallobase of the same length as parameres. Parameres simple, relatively broad, their sides attenuating sinuously to subapical hook. Parameral subbasal hook small, but the subapical hook is well developed. Median lobe lanceolate (Fig. 70). Female. Similar to male in habitus, but larger. Antenna shorter, and more slender. Differential diagnosis. E. similis sp. nov. is almost identical with E. jendeki in habitus. It differs from that species in the shorter last antennomere, sinuous inner side of paramere, and lanceolate median lobe of aedeagus. The short last antennomere and lanceolate median lobe distinguish E. similis sp. nov. also from another similar species E. siamensis sp. nov. Collection circumstances. Collected at light. Distribution. A species occurring in the upland area of northern Thailand and Laos. Etymology. The Latin word for “similar” refers to its similar appearance with other Eulichas species, in particular with E. jendeki.

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Eulichas uniformis (Pic, 1911) (Figs. 7, 31, 51, 71) Lichas uniformis Pic, 1911: 142 (original description, Bengale: Kurseong). Eulichas uniformis (Pic, 1911): Pic 1914: 11 (catalogue); Jäch 1995: 363 (lectotype designation, catalogue); Jäch & Hájek 2006: 455 (catalogue).

Type locality. “Bengale: Kurseong” [Kārsiyāng, West Bengal, India]. Type material. Lectotype ♂ (MNHN), designated by Jäch (1995): “Kurseong / Nord-Bengal / H.Rolle,Berlin,S.W.11 [black frame, printed] // Muséum Paris / Coll. M. Pic [yellow label, printed] // n sp [handwritten] // type [handwritten] // type [handwritten] // TYPE [red label, printed] // Lichas / uniformis Pic [handwritten] // LECTOTYPUS / Eulichas / uniformis PIC / des. M. Jäch 1995 [red label, handwritten]”. Additional material studied. 102 specimens — BHUTAN or INDIA: 8♂♂ 1♀, British Bootang, 1898, L. Durel (MNHN). INDIA: 1♀ , Assam (MNHN); 2♂♂ 1♀, Meghā laya, Khasia Hills [Khā si Hills], vii. [18]94 (MNHN); 1♀, Meghālaya, Jaintia Hills, Jowai, 25°27’N 92°12’E, 1350 m, 6.–8.vi.1996, E. Jendek & O. Šauša leg. (NHMW); 2♂♂, Meghālaya, W Garo Hills, Nokrek N.P., 25°29.6’N 90°19.5’E, 1100 m, 9.– 17.v.1996, E. Jendek & O. Šauša leg. (NHMW); 1♂, Sikkim, Rinchingpong [ca. 27°14’N 88°15’E], 1000 m, 22.iv.1987, Ch.J. Rai leg. (NHMB); 1♂, Sikkim, Buhuk Rain Forest, 1600 m, vi.2002, Malicky leg. (NMPC); 1♀, Uttar Pradesh, Allahābād [ca. 25°27’N 81°50’E], Bowring (BMNH); 2♀♀, West Bengal, Kārsiyāng [ca. 26°53’N 88°16’E], 1904, ex coll. Baron Moffaerts (BMNH); 1 ♂ , West Bengal, Darjeeling Distr., Bijanbari,800 m, 12.v.1975, W. Wittmer leg. (NHMB); 1♂, West Bengal, Darjeeling Distr., -Jhepi, 17.v.1975, W. Wittmer leg. (NHMB). MYANMAR: 8♂♂ 1♀, 25 km E Putao, Nan Sa Bon vill. env., 800 m, 6.–9.v.1998, S. Murzin & V. Siniaev leg. (NHMW, NMPC); 5♂♂, 50 km E Putao, Nan Thi env., 11.–16.v.1998, S. Murzin & V. Siniaev leg. (NMPC); 3♂♂ 1♀, 65 km NW Putao, Zi Yar Dam, 1250 m, 18.–21.v.1998, S. Murzin & V. Siniaev leg. (NMPC). NEPAL: 1♂, Indrawati Khola Saretar, 1700 m, 25./26.iv.1962, G. Ebert leg. (ZSMC); 2♂♂, Sun Khosi Tal [= valley], 2150 m, 2.v.1962, G. Ebert leg. (ZSMC); 4♂♂, Tampa Khosi Tal [= valley], 2600 m, 10.v.1962, G. Ebert leg. (ZSMC); 1♂ 3♀♀, Liukhu Khola Tal [= valley], 1700 m, 4.vi.1962, G. Ebert leg. (NMPC, ZSMC); 1♂, Bagmati distr., 4 km S of Tarang Marang, 900 m, 28.iv.1981, I. Löbl & A. Smetana leg. (MNHG); 1♀, Kathmandu distr., Kakani [ca. 27°49’N 85°14‘E], at light, 23.v.–20.vi.1983, M.G. Allen leg. (BMNH); 2♂♂ 1♀, Janakpur distr., Jiri-Shivalaya (Khimti-Khola), 1800–2500 m, 11.–12.vi.1987, C. Holzschuh leg. (NHMW); 1♂, same label data, but Ch.J. Rai leg. (NHMB); 3♂♂ 1♀, Janakpur distr., TambaKoshi-Khola, SE Charikot, 900–1200 m, 16.–25.vi.1987, C. Holzschuh leg. (NHMW); 5♂♂, same label data, but Ch.J. Rai leg. (NHMB, NMPC); 1 ♂ , Kathmandu distr., B ā l ā ju, [ca. 27°44’N 85°18’E], 1400 m, 18.v.1989, M. Brancucci leg. (NHMB); 1♂, Bagmati distr., Trisuli Bazar-Samri, Bhanjyang, 540–1900 m, 8.vi.1993, Probst leg. (NHMW); 4 ♂♂ 4 ♀♀ , Dolakha Distr., Suridhoban, 1050 m, 27.–28.v.2000, W. Schawaller leg. (NMPC, SMNS); 3♂♂ 10♀♀, Dolakha Distr., lower Khare Khola, 1400 m, 29.v.2000, W. Schawaller leg. (NMPC, SMNS); 3♂♂ 4♀♀, same label data, but 1200 m, 3.–4.vi.2000, (NMPC, SMNS); 1♂ 2♀♀, Dolakha Distr., Bazar bridge bellow Amatal, 1660 m, 8.vi.2000, W. Schawaller leg. (NMPC, SMNS); 1♀, Dolakha distr., lower Amata Khola, 1700 m, 9.vi.2000, W. Schawaller leg. (SMNS). Doubtful localisation: 1 ♂ , Inde Anglaise [India], Trichinopoly [Tiruchchir ā ppalli, Tamil Nadu], P. Newton leg. (NMPC). Description. Habitus elongate, fusiform. Body colouring from brownish-red to brown-blackish. Pale part of setation consists of recumbent yellow or greyish setae covering uniformly body surface, or forming indistinct ocellations on elytra (Fig. 7). Measurements. Males: 16–19 mm; females: 19–25 mm. Head punctation consists of irregularly distributed sparse moderately large setigerous punctures. Antenna relatively slender, last antennomere nearly rectangular, 2.67–3.25 times as long as wide (Fig. 31), its ventral side smooth. Pronotum trapezoidal, ca. 1.84–1.97 times as wide as long. Sides almost straight, with more or less



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distinct obtuse angle behind the middle (Fig. 51). Pronotum with two oval shallow depressions on the disc, and in some specimens with indistinct depressions also medially along anterior and posterior margin. Punctation consists of sparse moderately coarse setigerous punctures on the disc, which become slightly coarser and denser laterally. Elytra with numerous longitudinal rows of moderately large punctures, and very fine interstitial setigerous punctures. Ventral part with fine punctures, which are sparse medially and become larger and denser laterally. Last abdominal ventrite laterally almost regularly rounded to apex. Male. Aedeagus with phallobase of the same length as parameres. Parameres simple, relatively slender, slightly attenuating before subapical hook. Parameral subbasal hook small, but the subapical hook is well developed. Median lobe narrowly lanceolate (Fig. 71). Female. Similar to male in habitus, but larger. Antenna shorter, and more slender. Collection circumstances. Collected at light. Distribution. The species is known from upland area of Nepal, northern India (Uttar Pradesh, Sikkim, West Bengal, Meghālaya, Assam) and northern Myanmar (Kachin). The occurrence in Bhutan needs to be verified by precisely localised specimens. Eulichas sikkimensis species complex The species complex contains ten species distributed from Nepal and northern India, through continental south-eastern Asia, to Greater Sunda Islands and the Philippines. The highest diversity is reached in the Malay Peninsula. Species included in the complex are characterised by the reduction of the parameral subbasal hook.

Eulichas baeri (Fairmaire, 1898) (Figs. 8–9, 32, 52, 72) Lichas baeri Fairmaire, 1898: 388 (original description, Luzon). Eulichas baeri (Fairmaire, 1898): Pic 1914: 11 (catalogue); Jäch 1995: 363 (catalogue). Eulichas baeri var. innotatus Pic, 1924: 31 (original description, Luzon); Jäch 1995: 363 (catalogue). syn. nov.

Type locality. “Luzon” [Philippines] (E. baeri), “Luzon” [Philippines] (E. baeri innotatus). Type material. Lichas baeri: not studied, no syntype was found in Fairmaire’s collection (MNHN). Eulichas baeri var. innotatus: Lectotype ♂ (MNHN), by present designation: “Mt. Makiling / Luzon, Baker [printed] // 21160 [handwritten] // L. Baeri / Frm. var. [handwritten] // Muséum Paris / Coll. M. Pic [printed] / / TYPE [red label, printed] // v. innotatus / mihi [handwritten] // LECTOTYPUS / EULICHAS / baeri / var. innotatus Pic, 1924 / Jiří Hájek des. 2002 [red label, printed] // EULICHAS (s. str.) / baeri (Fairmaire, 1898) / Jiří Hájek det. 2002 [printed]”. Number of syntypes unknown. I designate a lectotype to fix the identity of this species, as available taxonomic works do not allow an unambiguous identification of specimens. Additional material studied. 155 specimens — BRUNEI: 1 ♂ , Bukit Teraja, Labi [ca. 04°24’N 114°27’E], Dipterocarp. forest, light trap, 60 m, 2.viii.1979, S.L. Sutton leg. (BMNH); 2♂♂ 1♀, Temburong, Kuala Belalong F.S.C., mixed Dipterocarp forest, 60–300 m, 16.–20.iv.1993, E. Heiss leg. (NHMW). INDONESIA: 1♀, Kalimantan Timur, Boven, Mahakkam, 1894, Dr. Nieuwenhuis exped. (RMNH); 1 ♂, Kalimantan Tengah, Busang / Rekut confluence, 0°03’S 113°59’E, light trap, M.J.D. Brendell & H. Mendel leg. (BMNH); 1♂, Kalimantan Timur, Sanga Sanga, H.D. Jensen leg. (BMNH). MALAYSIA: SABAH: 4♂♂ 7♀♀ , Kinabalu (HNHM, IRSNB, MNHN, SMTD); 1♂ 1 ♀, Mount Kinabalu (NHMB); 2♂♂ 4 ♀♀, same label data, but v.–viii.1903, J. Waterstradt (MNHN); 1♀, Kinabalu NP, Head Q., 06°00’N, 116°32’E, 1600 m, nearly undisturbed montane evergreen rain forest, 15.v.1987, J. Huisman leg. (RMNH); 2♀♀, Kinabalu Nat. Park, Sayap, 1000 m, 25.–29.xi.1996, W. Schawaller leg. (NMPC, SMNS); 3♀♀, same label data, but 6°10’N 116°34’E, primary forest edge, 950 m, at light, 8.iii.2001, J.P. Duffels & M.A. Schouten leg. (ZMAN); 1♀,

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Crocker Range, 16 miles NW of Keningau, 4.iv.1982, S. Nagai leg. (EUMC); 4♂♂ 8♀♀ , Crocker Range Mts., 50 km E Kota Kinabalu, Gunung Emas, 16.–27.iv.1993, I. Jeniš leg. (NHMW); 1♂ same label data, but 500–1900 m, 5.–21.v.1995, J. Stolarczyk leg. (NMPC); 1♂ 2♀♀, Banjaran Crocker Mts., 16 km SW Gunung Alab, 790–850 m, 4.–9.v.1996, M. Štrba & R. Hergovits leg. (NHMW); 1♀, 105 km S Beaufort, Long Pa Sia area, airstrip Long Pasia, 04°24’N, 115°43’E, 1000 m, at light, 15.iv.1987, v. Tol & Huisman leg. (RMNH); 1♂, 65 km W Lahad Datu, Sg. Purut camp, 04°57’N, 117°45’E, 250 m, at light, 26.x.1987, J. Huisman & R. de Jong leg. (RMNH); 1♂, 15 km W of Lahad Datu, Danum valley, light trap, 7.xii.1989, M.J. & J.P. Duffels leg. (ZMAN); 1♂, 70 km W Lahad Datu, Danum valley, Next to bridge Nature Trail, 150 m, rainforest along narrow stream, at light, M.J. & J.P. Duffels leg. (NMPC); 1♀, same label data, but 13.xii.1989 (ZMAN); Poling, near Ranau, 29.iv.1980, M. & A. Sakai leg. (EUMC); 1♂, Ranau Distr., Poring Hot Spring, Poring Lodge, 06°02’35’’N, 116°42’19’’E, ca. 650 m, 14.–16.v.2002, T. Kothe leg. (ZSMC); 1♀, West Coast zone, 12 km NNE Ranau, Poring Hot Springs, 06°04’N, 116°42’E, 550 m, at light, 9.–10.xi.1987, J. Huisman & R. de Jong leg. (RMNH); 2 ♂♂, Poring Hot Spring, 450–600 m, 9.–11.iii.2007, W. Schawaller leg. (SMNS); 1♂, 80 km E of Ranau, Telpid, 800 m, xii.1978, S. Nagai leg. (EUMC); 1♂ 1♀, Tawau [ca. 04°15’N 117°52’E], 20.iii.2006 (NMPC). SARAWAK: 2♀♀, 4th Division, Gunung Mulu National Park, at light, iii. 1978, J.D. Holloway et al. leg. (BMNH); 3♂♂ 1♀, same label data, but 150–200 m, v.–viii.1978, P.M. Hammond & J.E. Marshall leg. (BMNH); 1♂ , Rumah Kabau anak muggot, Ng Sebong Baleh, 25 km E Kapit, iii.1994, J. Kodada leg. (NHMW); 1♂, Kapit distr., Rumah Ugap vill., Sut. riv., 3.–9.iii.1994, S. Bílý leg. (NMPC); 1♂, Kapit distr., Sebong, Baleh river, 9.–21.iii.1994, J. Horák leg. (NHMW). PHILIPPINES: LEYTE: 1♂ (NMPC); 2♀♀, 22.viii.2003, J.L. Boundant leg. (MCSN); 1♂, Cabalian [= San Juan, ca. 10°16’N 125°09’E], coll. W. Schultze (SMTD); 1♂, Mt. Balocaue, 700 m, local collector leg. (NMPC). LUZON: 1♂, Mt. Isarog (SMTD); 1♀, Pangasinan [most probably = Pangasanan, ca. 14°58’N 120°42’E], coll. Benesh (SMTS); 2♀♀, Sorsogon [ca. 12°58’N 124°00’E] (NHMB); 1♀, Upper Bintacan, Isabela, 30.iv.1983, S. Osada leg. (EUMC); 1♂, Nueva Viscaya, Sta Fe, datton Pass, 900 m, at light, 3.–8.vi.1985, R.A. Müller leg. (RMNH); 2♂♂ 1♀, Nueva Vizcaya, vi.2003, I.A. Lumawig leg. (MCSN, NMPC). MINDANAO: 1♀, coll. W. Schultze (SMTD); 1♂, Davao [ca. 07°04’N 125°36’E], Dr. Platen (MNHN); 1♂, Davao, iii.1968 (EUMC); 1♀, Davao, 5.xi.1991 (MCSN); 1♀, Davao prov., Baracatan, Mt. Talomo, 1100 m, 3.–6.viii.1985, M. Owada leg. (EUMC); 1♀, Upper Barakatan, Apo Range, 1100 m, 17.viii.1985, M. Tomokuni leg. (EUMC); 1♀ , Baracatan, Eagle Centre, N slopes of Mt. Apo, 1100 m, 5.viii.1985, M. Sakai leg. (EUMC); 1♀, Agko, Mt. Apo [ca. 07°01’N 125°16’E], 1000 m, 7.x.1978, S. Nagai leg. (EUMC); 2♀♀, same label data, but 9.iii.1979 (EUMC); 1♂, Mt. Apo, 8.–10.i.1981, Natives leg. (EUMC); 2♂♂, Mt. Apo, iv.2005, I.A. Lumawig leg. (MCSN); 1♂, Lindaban [ca. 8°17’N 124°48’E], Bukindon, W. Schultze leg. (SMTD); 1♂, Manila, coll. Schwarz (DEIC); 1♀, Surigao prov., W. Boettiker leg. (SMTD); 1♂, Surigao [ca. 9°47’N 125°28’E], Baker leg. (SMTD); 1♂, Surigao, W. Schultze leg. (SMTD); 2♂♂ 2♀♀, Surigao, 15.v.1915 (NMPC, ZMHB); 1♀, Tudaya-Sibulan, 30.vii.1970, T. Okadome leg. (EUMC); 1♀, Gasy, Maitum, S Cotabato, 700 m, 12.viii.1985, M. Sakai leg. (EUMC); 2♀♀, South Cotabato, Parker Mts., Salacale, 600–900 m, at light, 2.–12.iv.1985, R.A.Müller leg. (RMNH); 1♂, South Cotabato, Koronadal Barrio 8, 100–300 m, 12.–14.vii.1986, R.A. Müller leg. (RMNH); 1♂ 1♀, Bagong Silang, 30 km NW Maramag, 1700 m, 13.–17.v.1996, BOLM leg. (NMPC, SMTD); 7♂♂ 3♀♀ , Davao Oriental, Boston, Agtuuganon Mt., Camp 55, 1020 m, 29.-v.-7.vi.1997, Müller, Buenafe & Gorost. leg. (NMPC, RMNH); 1♂, Surigao del Sur Lianga, 8 km W Diatagon, 08°42’N 126°05’E, 200 m, 3.–7.vii.2005, J.H. Lourens leg. (BMNH); 1♂ 2♀♀, Agusan Sur, 10 km SE Trento Sta Maria, 08°01.615’N, 126°12.322’E, 185 m, 27.–28.iv.2008, J.H. Lourens leg. (IRSNB); 1 ♂ , Davao Oriental, Aliwagwag Primary forest, 07°43.667’N, 126°17.304’E, 90 m, 30.iv.–1.v.2008, J.H. Lourens leg. (IRSNB). MINDORO: 3♂♂, Abra de Ilog [ca. 13°27’N 120°43’E], W. Schultze leg. (NMPC, SMTD). NEGROS: 1♂, vi.1991 (MCSN); 1♂, v.1996 (NMPC). PALAWAN: 2♂♂ 1♀, Cleopatra Needle Nat. Park, Tanabag river valley, 300 m, 20.–22.xii.1990, BOLM leg. (NHMB, NMPC). SAMAR: 1♂, Baker leg. (MNHN); SIBUYAN: 1♂ 1♀, Romblon, vii.1985 (EUMC); 12♂♂ 2♀♀, Romblon prov., Magdiwang, Pawala river, Tampayan Camp, Ga-ong area, 70–400 m, at light, 19.iii.–7.iv.1987, R.A. Müller leg. (NMPC, RMNH). Description. Habitus elongate, fusiform. Body colouring from brownish-red to brown. Pale part of REVISION OF EULICHAS II


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setation consists of recumbent yellowish or greyish setae covering uniformely body surface, or forming more or less distinct ocellations on elytra (Figs. 8–9). Measurements. Males: 13–24 mm; females: 20–26 mm. Head punctation consists of sparse, irregularly distributed moderately large setigerous punctures. Antenna slender, last antennomere rather oval, 2.57–3.58 times as long as wide (Fig. 32), its ventral side smooth. Pronotum trapezoidal, ca. 1.71–1.93 times as wide as long. Sides of rather variable shape. In some specimens skewed and straight in anterior half, with indistinct angle near the middle, and hind angles distinctly prominent (Fig. 52). On the other hand almost regularly rounded in some specimens. The disc with two indistinct rounded depressions. Punctation consists of sparse moderately coarse setigerous punctures on the disc, which become slightly coarser and denser laterally. Elytra with numerous longitudinal rows of moderately large punctures, and very fine interstitial setigerous punctures. Ventral part with fine punctures, which are sparse medially and become larger and denser laterally. Last abdominal ventrite laterally almost regularly rounded to apex. Male. Aedeagus with phallobase of the same length as parameres. Parameres simple, relatively broad, subparallel. Parameral subbasal hook is missing, but the subapical hook is developed. Median lobe narrowly lanceolate (Fig. 72). Female. Similar to male in habitus, but larger. Antenna shorter, and more slender. Remarks. So far the only Eulichas species known from the Philippines, thus I was able to identify it, although no syntype was found/recognised in Fairmaire’s collection (MNHN). A variable species. The variety innotatus represents only indistinctly ocellated specimens. It is without any doubt conspecific with typically coloured specimens, and is therefore junior subjective synonym of E. baeri. However, populations from different Philippine islands, as well as from Borneo, differ in body size, shape of last antennomere and pronotum. In particular, specimens from Borneo have usually more rounded sides of pronotum and more distinct ocellations on elytra, and may represent a separate subspecies. However, as the same characters can also be observed in some specimens from different Philippine Islands, I refrain from its formal description. Collection circumstances. Collected at light. Distribution. Known from the Greater Sunda Island of Borneo – Brunei, Indonesia (Kalimantan), Malaysia (Sabah, Sarawak), and the Philippines (Leyte, Luzon, Mindanao, Mindoro, Negros, Palawan, Samar).

Eulichas gigantea (Fairmaire, 1891) (Fig. 10) Lichas giganteus Fairmaire, 1891: CXXVIII (orig. description, Kaschmir). Eulichas gigantea (Fairmaire, 1891): Pic 1914: 11 (catalogue); Jäch 1995: 363 (catalogue); Jäch & Hájek 2006: 455 (catalogue).

Type locality. “Kashmir” [Kulu, Himāchal Pradesh, India]. Type material. Holotype ♀ (MNHN): “Kulu [printed] // Lichas / giganteus / Fairm. K. [handwitten, Fairmaire] // ex Musaeo / W. Rotschild / 1899 [white label with black frame, printed] // TYPE [red label with black frame, printed] // MUSÉUM PARIS / 1952 / COLL. R. OBERTHUR [yellow label, printed] // Holotypus / LICHAS / giganteus Fairmaire, 1891 / Jiří Hájek det. 2005 [red label, printed] // EULICHAS / gigantea (Fairmaire, 1891) / Jiří Hájek det. 2005 [printed]”. Description of holotype female. Habitus elongate, fusiform. Body colouring brownish-red. Setation consists of recumbent sparse yellowish setae forming very indistinct ocellations on pronotum and elytra (Fig. 10). [The type specimens is worn dorsally, originally the body pubescence was probably more distinct.] Measurements. Female: 31 mm.

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HÁJEK

Head punctation consists of irregularly distributed moderately large setigerous punctures. Antenna short, last antennomere elongated, ca. 2.8 times as long as wide. Pronotum transverse, twice as wide as long. Lateral sides regularly rounded. The disc convex. Punctation consists of moderately large setigerous punctures on the disc, which become coarser and denser laterally. Elytra with numerous longitudinal rows of large setigerous punctures, and interstitial very fine punctures. Ventral part uniformly densely punctured with fine punctures. Last abdominal ventrite laterally almost regularly rounded to apex. Male. Unknown. Remarks. Without a male available for study, the classification of E. gigantea in E. sikkimensis species complex is problematic. The only reason for its preliminary classification here is the similarity in habitus with E. robusta sp. nov. Distribution. So far known only from the type locality in Himāchal Pradesh, India.

Eulichas incisicollis Pic, 1933 (Figs. 11–12, 33, 53, 73) Eulichas incisicollis Pic, 1933: 22 (original description, Archipel Malais); Jäch 1995: 363 (catalogue); Ivie & Jäch 2002: 166 (correction of the type locality).

Type locality. “Archipel Malais: Isle” [“Camerons Island”, = Cameron Highlands, Malaysia]. Type material. Lectotype ♀ (BMNH), by present designation: “Type [printed, round label with red frame] // Pres. by / Imp.Inst.Ent. / Brit. Mus. / 1932-412 [printed] // G. 837 [handwritten vertically along the left side of the label] / Malaya [printed] / Camerons / Island. / 2.4.1930 [handwritten] / Entom. Div. / Agric. Dept. [printed] // Eulichas / incisicollis / n sp / (desiré) [Pic’s handwriten] // LECTOTYPE / EULICHAS (s. str.) / incisicollis Pic, 1933 / Jiř í Hájek det. 2009 [printed, red label]”. Number of syntypes unknown. I designate a lectotype to fix the identity of this species. There are several other species of Eulichas occurring in Cameron Highlands, and available taxonomic works do not allow an unambiguous identification of specimens. Additional material studied. 16 specimens — MALAYSIA: 1♂, Pahang, Cameron Highlands, 4800– 5500 ft., 8.vi.1935, H.M. Pendlebury leg. (MNHN); 1♀, Pahang, Cameron Highlands, Tanah Rata, 4800 ft., 12.ii.1926 (BMNH); 1♀, same label data, but 20.vii.1938, H.M. Pendlebury leg. (BMNH); 1♀, same label data, but 25.ii.1970, S. Suzuki leg. (NMPC); 1♀, same label data, but 4.iii.1970, S. Suzuki leg. (EUMC); 1♂, same label data, but 1.-6-iv.1990, A. Riedel leg. (SMNS); 1♂, same label data, but 1600 m, 11.–27.ii.2000, J. Horák leg. (NMPC); 1♂, same label data, but 1400 m, v.2000, L. Černý leg. (NMPC); 1♀, same label data, but 1500–1700 m, 1.–13.ii.2003, M. Oboř il leg. (NHMW); 1 ♀ , same label data, but P. Pacholátko leg. (NMPC); 1♂, same label data, but i.2006, P. Viktora leg. (NMPC); 1♂, Cameron Highlands, i.1985, S. Nagai leg. (EUMC); 1♂, Pahang, Cameron Highlands, 2 km S Tanah Rata, on Tapah Road, montane rainforest, at light, 29.iii.1993, O. Merkl & I. Szikossy leg. (HNHM); 1♀ , Pahang, Kampung Kuala Boh, 4°27.9’N 101°34.8’E, 850–1050 m, 26.iii.–3.iv.2001, M. Štrba & R. Hergovits leg. (NMPC); 1♂ 1♀, [Perak], Taiping [ca. 04°51’N 100°43’E], 6.v.1981, ex coll. S. Riese (MCSN). Description. Habitus elongate, fusiform. Body colouring from brownish-red to brown. Pale part of setation consists of recumbent yellow or whitish setae forming indistinct ocellations on pronotum, and very characteristic ocellations on elytra (Fig. 11–12). Measurements. Males: 21–28 mm; females: 33–37 mm. Head punctation consists of sparse, irregularly distributed moderately large setigerous punctures. Antenna slender, short compared to other species – reaching only the middle of body length. Last antennomere narrowly oval, 2.70–3.15 times as long as wide (Fig. 33), its ventral side smooth. Pronotum trapezoidal, ca. 1.95–2.11 times as wide as long. Sides skewed and straight in anterior two



27

thirds, then with obtuse angle, distinct incision subbasally, and prominent hind angles (Fig. 53). Pronotum with two rounded shallow depressions on the disc, and indistinct depressions medially also along anterior and posterior margin. Punctation consists of sparse regularly distributed setigerous punctures. Elytra with numerous longitudinal rows of moderately large punctures, and very fine interstitial setigerous punctures. Ventral part with fine punctures, which are sparse medially and become larger and denser laterally. Last abdominal ventrite laterally regularly rounded to apex. Male. Aedeagus with phallobase shorter than parameres. Parameres simple, slender, their inner side attenuating regularly to subapical hook. Parameral subbasal hook is missing, but the subapical hook is developed. Median lobe slender, rather subparallel (Fig. 73). Female. Distinctly larger than male. Antenna shorter, and more slender. Lateral incision of pronotum more distinct. Collection circumstances. Collected at light. Distribution. So far known only from the Cameron Highlands on the Malay Peninsula.

Eulichas oborili Hájek, sp. nov. (Figs. 13, 34, 54, 74) Type locality. Thailand, Nan prov., Ban Bo Klua env. Type material. 3 specimens — Holotype ♂ (NMPC), labelled: “THAILAND 13.–26.V.2002 / NAN Prov. / BAN BO KLUA env. / P. Průdek & M. Obořil leg. [printed]”. Paratypes: 2♀♀ (nos. 1–2), same data as holotype (NMPC). Description. Habitus elongate, fusiform. Body colouring from brown to brown-blackish. Pale part of setation consists of recumbent yellowish grey setae forming characteristic ocellations on pronotum, elytra and abdominal sternites (Fig. 13). Measurements. Male: 23 mm; females: 27–28 mm. Head punctation consists of sparse, irregularly distributed moderately large setigerous punctures. Antenna long, slender, last antennomere nearly rectangular with apical elongation, ca. 3.65 times as long as wide (Fig. 34), its ventral side smooth. Pronotum trapezoidal, ca. 1.81–1.90 times as wide as long. Sides almost regularly rounded (Fig. 54). The disc with two oval shallow depressions. Punctation consists of sparse moderately coarse setigerous punctures on the disc, which become slightly coarser and denser laterally. Elytra with numerous longitudinal rows of moderately large punctures, and very fine interstitial setigerous punctures. Ventral part with fine punctures, which are sparse medially and become larger and denser laterally. Last abdominal ventrite laterally regularly rounded to apex. Male. Aedeagus with phallobase of the same length as parameres. Parameres simple, slender, parallel sided, with relatively short apical part. Their subbasal hook is missing, but the subapical hook is well developed and prominent. Median lobe lanceolate, slightly exceeding parameres (Fig. 74). Female. Similar to male in habitus, but larger. Antenna shorter, and more slender. Differential diagnosis. In habitus, E. oborili sp. nov. is similar to E. siamensis sp. nov. and E. similis sp. nov. of the E. uniformis complex, but it is more elongate and subparallel. It could be easily recognised by the shape of aedeagus. E. oborili sp. nov. has slender, parallel sided parameres without subbasal hook, and distinctly lanceolate median lobe exceeding the apex of parameres. Collection circumstances. Collected under the bark of tree with fermenting sap, near a stream (M. Obořil pers. comm., 2005). Distribution. So far known only from the type locality in northern Thailand. Etymology. The new species is dedicated to one of its collectors, my friend Martin Obořil (Brno, Czech

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HÁJEK

Republic), a specialist on Buprestidae.

Eulichas robusta Hájek, sp. nov. (Figs. 14, 35, 55, 75) Type locality. Malaysia, Pahang, Bukit Fraser, ca. 03°43’N 101°44’E, 1500 m. Type material. 130 specimens — Holotype ♂ (NMPC), labelled: “Malaysia 1500m / Bukit Fraser / 22.– 28. 11. 1998 / R.Kocina lgt. [printed]”. Paratypes: 2♂♂ (nos. 1–2) 1♀ (no. 3), same label data as holotype (NMPC); 1♂ (no. 4) 2♀♀ (nos. 5–6), “MALAYSIA 24.11.[19]98 / Bukit fraser hill / H. Habal lgt. [printed]” (NMPC); 1♂ (no. 7), “MALAYSIA / BUKIT FRASER / 1. – 5. 5. 2003 / B. Makovský lgt. [printed]” (NMPC); 5♂♂ (nos. 8–12), “MALAYSIA / BUKIT FRASER / 25. – 31.3 2004 / M. Machytka lgt. [printed]” (NMPC); 6♂♂ (nos. 13–18) 1♀ (no. 19), same label data, but “B. Makovský lgt. [printed]” (NMPC); 1♀ (no. 20), “MALAYSIA / BUKIT FRASER / 8. – 14. 4. 2008 / B. Makovský lgt. [printed]” (NMPC); 1♂ (no. 21), “W.Malaysia: 1140 m / Fraser’s Hill, / Jeriau Road / 5–12.viii.1986 [printed] // G.S.Robinson / BM 1986–299 [printed]” (BMNH); 1♀ (no. 22), “W.Malaysia: Pahang / Fraser’s Hill,ca 1300m /17–21.3.[19]93, light trap / Löbl&Calame, #14 [printed]” (MNHG); 1♂ (no. 23), “WEST MALAYSIA, PAHANG / 7.–10. 1. 1995 / FRASER’S HILL, 1500 m / 110 km N KUALA LUMPUR / lgt.S.BECVAR j. & s [printed]” (BMNH); 6♂♂ (nos. 24–29), “Malaysia: 10.V. / Fraser’s Hill / S.Snäll lgt. 1996 [printed]” (SSCT); 5♂♂ (nos. 30–34), “Malaysia Pahang Fraser’s / Hill 1200m 23–25.3 1992 light / B Gustafsson, H et H Hippa / G Sellerholm [printed]” (NHRS); 1♀ (no. 35), “MALAYSIA - Pahang / Banjaran Benom / Lata Jarom / 6.–8.3.1997 / Ivo Jeniš leg. [printed]” (NMPC); 1♂ (no. 36), same label data, but “Oliver Dulík leg. [printed]” (NMPC); 2♂♂ (nos. 37–38) 2♀♀ (nos. 39–40), “MALAYSIA iii.1999 / Pahang & Kelantan prov. / Cameron Highlands. / lgt. local collector [printed]” (NMPC); 1♀ (no. 41), “Weiden / Opf. [printed] / Cameron / Highlands [handwritten] / [on side:] STRÖHLE [printed]” (VKCZ); 1♂ (no. 42), “Malaysia / Pahang / Cameron Hids / 1000–1600ft / III–VII/1978 / C.M.Brandstetter [printed]” (SMNS); 1♂ (no. 43), “MALAYSIA / Tanah Rata / V. 1987. / leg. B. Molnár [handwritten]” (HNHM); 1♂ (no. 44), “MALAYSIA - Perak / Cameron Highlands / Tanah Rata / 13. – 16.3.1997 / Ivo Jeniš leg. [printed]” (NMPC); 2♂♂ (nos. 45–46) 1♀ (no. 47), “MALAYSIA W / CAMERON highl. / TANAH RATA env. / 23.–26.3.2000 / Lgt. M. SNÍŽEK [printed]” (NMPC); 1♂ (no. 48) 1 ♀ (no. 49), “MALAYSIA West, PAHANG / Cameron Highlands / TANAH RATA, 1200–1500m / 3.ii.– 19.ii. 2005 / Cechovsky Petr lgt [printed]” (VKCZ); 1♂ (no. 50), “MALAYSIA / TANAH RATA / 2. – 8. 4. 2008 / B.MAKOVSKÝ Lgt. [printed]” (NMPC); 1♂ (no. 51), “28 VII 1991 U.V 750. / Rte TAPAH – TANAH RATA / PK 40 CAMERON HIGHLANDS / MALAYA Leg. J.HAXAIRE [printed]” (NHMW); 1♂ (no. 52), “W Malaysia - Perak / road Tapah - Ringlet / 10km S of Ringlet mt. 900 / 14–19.IV.1999 / leg. A. Ballerio [printed]” (NMPC); 3 ♂♂ (nos. 53–55), “MALAYSIA W / CAMERON Highlands / RINGLET env. / 15.4.2000 / LGT. M. SNÍŽEK [printed]” (NMPC); 1 ♂ (no. 56), “MALAYSIA - PERAK Distr. 20.– 23.XI.2000 / Cameron Highlands / RINGLET env. / F. & L. Kantnerovi leg. [printed]” (NMPC); 5♂♂ (nos. 57–61) 1 ♀ (no. 62), “MALAYSIA W., PERAK / 40km SE of IPOH, 900 m / Banjaran Tai Wangsa / RINGLET, 29.iii.–15.iv. / 2004 Čechovsky Petr lgt. [printed]” (NMPC, VKCZ); 1♂ (no. 63) 1♀ (no. 64), “MALAYSIA –W; PERAK, ~900m; / 40 km SE IPOH; 4°25’N 101°23’E / Cameron Highlands; RINGLET; / M. Ř íha leg. 25.iv.–5.v.2001 [printed]” (NMPC); 1 ♂ (no. 65), “MALAYSIA, Pahang distr. / Cameron Highlands, 2001 / KAMPUNG KUALA BOH vill. env. / 4°27’N, 101°34’E, 850–1050m / R. Hergovits leg. 26. III.–3. IV. [printed]” (NMPC); 1 ♂ (no. 66), “MALAYSIA, Pahang distr. / Cameron Highlands, / KAMPUNG KUALA BOH vill. env., / N 4°27,9’, E 101°34,8’, 850–1050m, / 26. III.–3. IV. 2001 / K. Bucsek leg. [printed]” (NMPC); 3♂♂ (nos. 67–69), “MALAYSIA / Cameron / Highlands [printed]” (ZSMC); 1♂ (no. 70), “W. Malaysia / Cameron / Highlands / 2000 ft. above / sea level. 18.2.[19]73 / C.C.Chua [handwritten] // Brit. Mus. / 1975-573 [handwritten]” (BMNH); 1♂ (no. 71), same label data, but “5.3.[19]75” (BMNH); 4♂♂ (nos. 72–75), “W – MALAYSIA / Cameron IV. / Highland 1987 [printed]” (NHMW); 6♂♂ (nos. 76–81) 2♀♀ (nos. 82–83), “Cameron Highlands / MALAYSIA / late April 1987 [printed]” (EUMC); 1♂ (no. 84), “Perak REVISION OF EULICHAS II


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[handwritten] / Fry Coll. / 1905-100. [printed]” (BMNH); 1♂ (no. 85), “64723 [handwritten] / Perak [printed] / Fry Coll. / 1905-100. [printed]” (BMNH); 1♂ (no. 86), “MALAYA [printed] / Perak / Gunong Kledang / 2500’ / 12 Feb 1959 [handwritten] / H. T. Pagden [printed] // C.I.E. COLL. / NO. [printed] 16488 [handwritten] // Eulichas sp. / ? subocellata / Fairm. [handwritten] / R.D.Pope det. 1959 [printed] // Pres by / Com inst Ent / B M 1959-499 [printed]” (BMNH); 1♂ (no. 87), W MALAYSIA, Perak distr., / 48 km NNE of IPOH, 2001 / FELDA LASAH vill. env., / 5°02’N, 101°12’E, 120–390 m / R. Hergovits leg. 13–21.III. [printed]” (NMPC); 1♂ (no. 88), “W MALAYSIA, Perak distr., / FELDA LASAH vill. env., 48 km NNE of IPOH / N 5° 02,3’, E 101° 12,3’, 120–390 m, / 13.–21.III.2001 / P. Šomody leg. [printed]” (NMPC); 1♀ (no. 89), “MALAYSIA - Perak / Banjaran Bintang / Maxwell Hill (Taiping) / 18. – 19.2.1997 / Ivo Jeniš leg. [printed]” (NMPC); 1♀ (no. 90), same label data, but “lgt. Oliver Ďulík [printed]” (VKCZ); 1♂ (no. 91), “MALAYSIA-Perak / Bukit Larut / 23.–25.2.2000 / K. Deneš jun. lgt. [printed]” (VKCZ); 3♂♂ (nos. 92–94), “WEST MALAYSIA, Perak / BUKIT LARUT (Maxwell Hill) / Taiping–Bintang Mts., (WGS84) / 04° 51’ 37’’N, 100° 47’ 56’’E / 9./13.7.2001, alt. 1100–1400m / lgt. R.Fouqué & H.Barlová [printed]” (NMPC); 2♂♂ (nos. 95–96), “Malaysia, Perak / Taiping, Bukit Larut 1036 m / 16.–17. 2. 1998 / V. Tichý lgt. [printed]” (NKME); 1♂ (no. 97), “Taiping / Malaysia / 3.1975 [handwritten] // Eulichas / funebris / Westwood. / det. Stig Lundberg [handwritten]” (NHRS); 1♂ (no. 98), “Malaysia / Taiping / May 1987 [handwritten]” (MCSN); 2♀♀ (nos. 99–100), “MALAYSIA / Taiping / X.1979 [handwritten]” (MSCN); 1♀ (no. 101), “TaipingMalaysia / VI-1982 [handwritten]” (MSCN); 6 ♀♀ (nos. 102–107), “MALAYSIA / Taiping / V.1987 [handwritten]” (MSCN, NMPC); 1♂ (no. 108), “MALAYSIA - Kelantan / Banjaran Titi Wangsa / Kampong Lawa env. / 24. – 26.2.1997 / Ďulík & Jeniš leg. [printed]” (NMPC); 6♂♂ (nos. 109–114) 1♀ (no. 115), “MALAYSIA 10.–16.iv.1999 / Kelantan prov. / Kampong Raja env. / lgt. Mir. Janalík [printed]” (NMPC, VKCZ); 1♂ (no. 116), same label data, but “lgt. Vít Kabourek [printed]” (VKCZ); 4 ♂♂ (nos. 117–120), “MALAY PENIN: [printed] / Selangor F.M.S. / Bukit Kutu 3457’ / April 1915 [handwritten] // Ex F.M.S. / Museum. / B.M. 1955-354. [printed]” (BMNH); 1♂ (no. 121), “MALAY PENIN: / Selangor. / Bukit Kutu [printed] / at light 3500 ft. / April 17th 1926 [handwritten] / H.M.Pendlebury. [printed] / [on reverse:] Ex. Coll: / F.M.S. Museum. [printed] // Ex F.M.S. / Museum. / B.M. 1955-354. [printed]” (BMNH); 2♂♂ (nos. 122– 123), same label data, but “April 20th” (BMNH); 1♂ (no. 124), “Malaysia [handwritten]” (MSCN); 1♂ (no. 125), “Pi Tam [Nakhon Si Thammarat prov., ca. 08°44’N 99°39’E] / S. Thailand / 8.VIII.1987 / M. Satô leg. [printed]” (EUMC); 1♂ (no. 126) 3♀♀ (nos. 127–129), “Saraburi [handwritten] / Thailand / leg. Sab. Steinke [printed] / [on side:] 1989 / XI. [handwritten]” (NMPC, SMNS). Additional material studied (doubtful localisation). 1♂, Philippines, Luzon, Sta Lucia Dolores Quezon, 500 m, at light, 25.vii.1991, J. Haxaire leg. (NHMW). Description. Habitus elongate, fusiform. Body colouring from brownish-red to brown. Pale part of setation consists of recumbent yellowish or yellow greyish setae forming indistinct ocellations on pronotum, and characteristic ocellations on elytra (Fig. 14). Measurements. Males: 26–33 mm (holotype 32 mm); females: 32–37 mm. Head punctation consists of sparse, irregularly distributed moderately large setigerous punctures. Antenna long, slender, last antennomere rectangular, with apical elongation, 3.19–3.63 times as long as wide (Fig. 35), its ventral side smooth. Pronotum trapezoidal, ca. 2.01–2.04 times as wide as long. Sides almost regularly rounded (Fig. 55). The disc with two indistinct rounded depressions. Punctation consists of sparse fine setigerous punctures on the disc, which become slightly coarser and denser laterally. Elytra with numerous longitudinal rows of moderately large punctures, and very fine interstitial setigerous punctures. Ventral part with fine punctures, which are sparse medially and become larger and denser laterally. Last abdominal ventrite laterally with distinct sinuation before apex. Male. Aedeagus with phallobase of the same length as parameres. Parameres simple, broad, slightly

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HÁJEK

attenuating to subapical hook. Their subbasal hook is missing, but the subapical hook is well developed and prominent. Median lobe lanceolate (Fig. 75). Female. Similar to male in habitus, but larger. Antenna shorter, and more slender. Differential diagnosis. Eulichas robusta sp. nov. can be easily recognised from other Malayan species based on its large body size, regularly rounded sides of pronotum, and predominantly by the characteristic shape of aedeagus: Broad parameres with robust apical part, and distinctly lanceolate median lobe. However, the new species is very similar to E. gigantea known only from the female holotype. The only usable chatacter to distinguish between these species is the shape of pronotum, which is more rounded laterally in E. gigantea. Nevertheless, both species occur nearly 4.000 km from each other, and I do not expect them to be conspecific. Collection circumstances. Collected at light. Distribution. Known from number of localities in Malay Peninsula (Kelantan, Pahang, Perak, Selangor), and two localities in southern Thailand (Nakhon Si Thammarat, Saraburi). Etymology. The Latin word for “robust” refers to generally large habitus of the species.

Eulichas sausai Hájek, sp. nov. (Figs. 15, 36, 56, 76) Type locality. Malaysia, Pahang, ca. 30 NE of Raub, Lata Lembik, ca. 03°56’N 101°38’E, 200–400 m. Type material. 6 specimens — Holotype ♂ (NMPC), labelled: “MALAYSIA, Pahang distr., / 30km NE RAUB, LATA LEMBIK / 3°56’N; 101°38’E, 200–400 m / 22.IV–1.V., 8–15.V.2002 / E. Jendek & O. Šauša leg. [printed]”. Paratypes: 1♂ (no. 1), “Malaysia–Taman Negara [Nat. Park] / Kuala Tahan [Pahang distr.] / 22 March 1988 / leg. G. Hangay [printed]” (HNHM); 1♂ (no. 2) 1♀ (no. 3), “N.O: Sumatra / Deli [= Medan, ca. 03°35’N 98°40’E] / L.Martin S. [yellow label, printed] // 86991 [handwritten]” (ZMHB); 2♂♂ (nos. 4–5). “Sumatra / Indragiri / J. Bouchard / 1905. [printed] // MUSÉUM PARIS / 1952 / COLL. R. OBERTHUR [yellow label, printed]” (MNHN). Additional material studied. 2 specimens — INDONESIA: SUMATRA: 1♂ NE Sumatra, R. Taylor (BMNH). MALAYSIA: 1♀, Sungkai reserve, 1.–10.vii.1918, ex F.M.S. Museum (BMNH). Description. Habitus elongate, fusiform. Body colouring from brownish-red to brown. Pale part of setation consists of recumbent yellowish or yellow greyish setae forming indistinct ocellations on pronotum, and characteristic ocellations on elytra (Fig. 15). Measurements. Males: 20–24 mm (holotype 24 mm); females: 25–30 mm. Head punctation consists of sparse, irregularly distributed moderately large setigerous punctures. Antenna long, slender, last antennomere rather oval, with apical elongation, 2.56–3.63 times as long as wide (Fig. 36), its ventral side smooth. Pronotum trapezoidal, ca. 1.92–1.97 times as wide as long. Sides almost regularly rounded (Fig. 56). The disc with two indistinct rounded depressions. Punctation consists of sparse fine setigerous punctures on the disc, which become slightly coarser and denser laterally. Elytra with numerous longitudinal rows of moderately large punctures, and very fine interstitial setigerous punctures. Ventral part with fine punctures, which are sparse medially and become larger and denser laterally. Last abdominal ventrite laterally regularly rounded to apex. Male. Aedeagus with phallobase shorter than parameres. Parameres simple, slender, slightly attenuating to subapical hook. Their subbasal hook is missing, but the subapical hook is well developed, slightly prominent. Median lobe slender, rather subparallel (Fig. 76). Female. Similar to male in habitus, but larger. Antenna shorter, and more slender. Differential diagnosis. E. sausai sp. nov. is similar to E. serricornis sp. nov. in habitus. However, it can be distinguished at first sight by the elytral ocellations forming more or less distinct transverse fasciae. In addition, E. sausai sp. nov. has less serrate antennomeres 3–10 and shorter antennomere 11, parameres REVISION OF EULICHAS II


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attenuating regularly to the subapical hook, and median lobe is subparallel, attenuating only in the apical third of its length. Collection circumstances. Specimens from Malaysia were collected at light. Distribution. The species is so far known only from several localities in Malay Peninsula and Sumatra Island. Etymology. The new species is dedicated to one of its collectors, Ondrej Šauša (Bratislava, Slovakia), a specialist on Elateridae.

Eulichas serricornis Hájek, sp. nov. (Figs. 16, 37, 57, 77, 85) Type locality. Malaysia, Perak, ca. 48 km NNE of Ipoh, Felda Lasah vill., ca. 05°02.3’N 101°12.3’E, 120– 390 m.

FIGURES 85–86. Habitat of Eulichas. 85—male of E. serricornis sp. nov. found sitting on vegetation at the locality “Batu 19”; 86—stream at the Batu 19 (Cameron Highlands, Perak, Malaysia), the habitat of E. serricornis sp. nov.

Type material. 60 specimens — Holotype ♂ (NMPC), labelled: “W MALAYSIA, Perak distr., / FELDA LASAH vill.env.,48 km NNE of IPOH / N 05° 02,3’, E 101° 12,3’, 120–390 m, / 13.–21.iii.2001 / M. Štrba leg. [printed]”. Paratypes: 3♂♂ (nos. 1–3), same label data as holotype (NMPC); 6♂♂ (nos. 4–9), same label data, but “P. Šomody leg.” (NMPC); 1♂ (no. 10), “PERAK, F. M. S. / Batang Padang / Jor Camp [printed] 1800 [handwritten] ft. [printed] / March 16th [handwritten] 1923 / H. M Pendlebury [printed] / [on reverse:] Ex Coll: / F. M. S. / Museums. // Ex F.M.S. / Museum. / B.M. 1955-354. [printed]” (BMNH); 1♂ (no. 11), same label data, but “June 4th 1923” (BMNH); 1♂ (no. 12), same label data, but “light Feb. 26th 1924” (BMNH); 1♂ (no. 13), same label data, but “Jan 26th 1925” (BMNH); 1♂ (no. 14), “PERAK F.M.S. / Batang Padang, / KUALA WOH / Mar: [printed] 20th [handwritten] 1940 [printed] / [on reverse:] Selangor Mus: / Collectors. [printed] // Ex F.M.S. / Museum. / B.M. 1955-354. [printed]” (BMNH); 1♂ (no. 15) 1♀ (no. 16), “TapahTanah R / W MALAYSIA / 1976 VII / Native coll. [printed]” (EUMC); 1♀ (no. 17), “12 II 2001 / 19miles Cameron / High Land Maley. / A. et R. ABE leg. [printed]” (EUMC); 3♂♂ (nos. 18–20), “MALAYSIA, PERAK / Cameron Highlands / BATU (= MILE) 19 vill. env. / 04°22.2’N, 101°20.0’E; 590 m / Jiří Hájek leg. 25.iv.–11.v.2009 [printed]” (NMPC); 1 ♂ (no. 21), “Cameron High / Malaysia / III. 1974 / Nat. coll.

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[handwritten]” (EUMC); 3♂♂ (nos. 22–24), “Cameron Highlands / MALAYSIA / late April 1987 [printed]” (EUMC); 1♂ (no. 25), “PAHANG, F. M. S. / Lubok Tamang [printed] / 3.500 [handwritten] ft. [printed] / June 11th [handwritten] 1923 / H. M. Pendlebury. [printed] / [on reverse:] Ex Coll / F. M. S. / Museums. [printed] // Ex F.M.S. / Museum. B.M. 1955-354 [printed]” (BMNH); 3♂♂ (nos. 26–28), “MALAYSIA - Pahang / Banjaran Benom / Lata Jarom / 6.–8.3.1997 / Ivo Jeniš leg. [printed]” (IJCM, NMPC); 1 ♂ (no. 29), “MALAYSIA Pahang Distr. / 30 km NE RAUB, Lata Lembik / (3°56’N, 101°38’E), 200–400 m / 22.IV.–1.V., 8.–15.V.2002 / E. Jendek & O. Šauša leg. [printed]” (NMPC); 1♂ (no. 30), “Malaysia, Pahang, Fraser’s / Hill, 1200 m 20.2 1991 light / B. Gustafsson, H et H Hippa / G. Sellerholm [printed]” (NHRS); 1♂ (no. 31), “TAMAN NEGARA NP / Kuala Juram, E. of Merapoh / 4°39’ N 102°08’ E / 13.III.1999 // MALAYSIA / Pahang / J.P. & M.J. Duffels, / M.Zaidi & M.Y.Ruslan // edge primary rainforest / (near dormitory) / at light [printed]” (ZMAN); 1♂ (no. 32), “MALAY PENIN: [printed] / Kelantan / Ledlad / 23.2. 1936 [handwritten] / / Ex F.M.S. / Museum. / B.M. 1955-354 [printed]” (BMNH); 4 ♂♂ (nos. 33–36), “MALAYSIA 10.– 16.iv.1999 / Kelantan prov. / Kampomg Raja env. / lgt. Mir. Janalík [printed]” (NMPC, VKCZ); 1♂ (no. 37), “MALAYSIA W., KELANTAN / Road between Kampong Raja / and Gua Musang, 1400–1700m / (Ladang Pandrak), 1–28.iv.2006 / 4°63’N-101°45’E – 4°88’N- / 101°95’E, Cechovsky Petr lgt. [printed]” (NMPC); 3♂♂ (nos. 38–40), “MALAYSIA / Cameron / Highlands [printed]” (ZSMC); 2♂♂ (nos. 41–42), “Taiping Malaysia / X-1980 // Dono S. Riese / 1984 [handwritten]” (MCSN); 3 ♂♂ (nos. 43–45), “Malaysia [handwritten]” (MCSN); 1♂ (nos. 46), “Malaysia / Ulu Gombak / Kuala Lumpur [printed] / July 21. 1971 [handwritten] / R. E. Parrott [printed]” (CNCO); 1♂ (no. 47) 1♀ (no. 48), same label data, but “July 22. 1971” (CNCO); 1♂ (no. 49), same label data, but “19-VIII-. 1971” (CNCO); 1♂ (no. 50), same label data, but “AUG 19. 1971” (CNCO); 1♀ (no. 51), same label data, but “AUG 21. 1971” (CNCO); 1♀ (no. 52), same label data, but “AUG 22. 1971” (CNCO); 4♂♂ (nos. 53–56) 3♀♀ (nos. 57–59), same label data, but “AUG 24. 1971” (CNCO, NMPC). Description. Habitus elongate, fusiform. Body colouring from brownish-red to brown. Pale part of setation consists of recumbent yellow greyish, or whitish setae forming indistinct ocellations on pronotum, and characteristic ocellations on elytra (Fig. 16). Measurements. Males: 20–24 mm (holotype 23 mm); females: 26–30 mm. Head punctation consists of sparse, irregularly distributed moderately large setigerous punctures. Antenna long, slender, antennomeres 3–10 distinctly serrate, last antennomere filiform, 4.01–5.10 times as long as wide (Fig. 37), its ventral side smooth. Pronotum trapezoidal, ca. 1.91–2.04 times as wide as long. Sides almost regularly rounded (Fig. 57). The disc with two oval shallow depressions. Punctation consists of sparse moderately coarse setigerous punctures on the disc, which become slightly coarser and denser laterally. Elytra with numerous longitudinal rows of moderately large punctures, and very fine interstitial setigerous punctures. Ventral part with fine punctures, which are sparse medially and become larger and denser laterally. Last abdominal ventrite laterally regularly rounded to apex. Male. Aedeagus with phallobase shorter than parameres. Parameres simple, parallel sided, slender – the most slender in the middle of their length. Their subbasal hook is missing, but the subapical hook is well developed. Median lobe slender, rather subparallel (Fig. 77). Female. Similar to male in habitus, but larger. Antenna shorter, and more slender. Differential diagnosis. The new species is similar to other Malayan species of Eulichas, in particular to E. robusta sp. nov. and E. sausai sp. nov. From the first mentioned species, it differs in its smaller body size; from all species, E. serricornis sp. nov. differs significantly in the distinctly serrate antennomeres 3–10, and long, nearly filiform antennomere 11. Collection circumstances. Most specimens were collected at light. At the locality “Batu 19” (Cameron Highlands, Perak, Malaysia), it was found sitting on vegetation near the riverbank (ca. 6–8 m wide, mostly with sandy bottom) in mountain rainforest, at the altitude ca. 590 m a.s.l. (Fig. 86). Distribution. So far known only from several localities in Kelantan, Pahang and Perak provinces in the REVISION OF EULICHAS II


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Malay Peninsula. Etymology. The name is derived from Latin words for “serrate” and “cornu” [= antenna], which refer to the characteristic shapes of antennomeres of the new species.

Eulichas sikkimensis (Pic, 1913) (Figs. 17, 38, 58, 78) Lychas sikkimensis Pic, 1913: 108 (original description, Indes: Sikkim). Lycas seillierei Pic, 1915: 7 (original description, Himalaya). Eulichas sikkimensis (Pic, 1913): Pic 1914: 11 (catalogue); Jäch 1995: 363 (lectotype designation, catalogue). Eulichas sellierei (Pic, 1915): Jäch 1995: 363 (lectotype designation, in synonymy).

Type locality. “Indes: Sikkim” (E. sikkimensis); “Himalaya” (E. sellierei). Type material. Lychas sikkimensis: Lectotype ♂ (MNHN), designated by Jäch (1995): “Sikkim [printed] // Muséum Paris / Coll. M. Pic [printed] // TYPE [red label, printed] // Lychas / sikkimensis / Pic / type [handwritten] // LECTOTYPUS / Eulichas / sikkimensis PIC / des. M. Jäch 1995 [red label, handwritten]”. Lycas seillierei: Lectotype ♂ (MNHN), designated by Jäch (1995): “Himalaya / oriental [printed] // Type [handwritten] // TYPE [red label, printed] // Muséum Paris / Coll. M. Pic [printed] // Seillieri / Pic [handwritten] // LECTOTYPUS / Eulichas / seillierei PIC / des. M. Jäch 1995 [red label, handwritten] // Eulichas / sikkimensis PIC / det. Jäch [handwritten]”. Paralectotype: 1♂ , same label data as lectotype (MNHN). Additional material studied. 35 specimens — BHUTAN or INDIA: 4♂♂ 5♀♀, British Bootang, 1898, L. Durel (MNHN). INDIA: 4♂♂ , Sikkim, Gopaldhara, H. Stevens leg. (BMNH); 1♂, Uttaranchal, West Bhatkot, Kumaon, 4000 ft. [ca. 1220 m], v.1920 (BMNH); 1♀, West Bengal, Kurseong env. [Kārsiyāng, ca. 26°53’N 88°16’E] (NMPC); 1♂ 2♀♀, same label data, but P. Braet leg. (IRSNB); 2♂♂ 4♀♀, West Bengal, Mungphu [most probably = Mangpu, ca. 26°58’N 88°21’E], coll. Atkinson (BMNH, NMPC); 1♂ , West Bengal, Darjeeling [distr.], Chibo Busty, 900 m, 24.iv.1986, Ch.J. Rai (NHMB); 1♀, West Bengal, Darjeeling [distr.], Kalimpong env., 600 m, 8.v.1981 (NMPC); 2♀♀ , West Bengal, Darjeeling [distr.], Kalimpong, Khangebung, 1400 m, 30.iv.1987, N. Dangal leg. (NHMW); 1♂, West Bengal, Darjeeling distr., Darjeeling, Sing Bari, 1600 m, 8.ix.1990, N. Dangal leg. (NHMW). NEPAL: 1 ♂ , Danda Pakhar, 1600–2500 m, 1.vi.1977, M. Brancucci leg. (NMPC); 2♂♂, Solu-Khumbo, Mt. Everest, Nuan Thala, 1800 m, 23.iii.1988, Pasang leg. (NHMW); 1♂ , Myagdi distr., Hille-Ghorepani, Dhawlagiri, 1600–2600 m, 10.vi.1986, C. Holzschuh leg. (NHMB); 1♂, Bagmati distr., Nuwākot, Pāti Bhanjyāng [ca. 27°51’N 85°27’E], 1900 m, 16.– 18.vi.1989, M. Brancucci leg. (NHMB). Doubtful localisation: 1♂, Japan (SMTD). Description. Habitus elongate, fusiform. Body colouring from brownish-red to brown. Pale part of setation consists of recumbent yellowish or greyish setae covering uniformly body surface, or forming very indistinct ocellations on elytra (Fig. 17). Measurements. Males: 16–19 mm; females: 20–22 mm. Head punctation consists of sparse moderately large setigerous punctures. Antenna slender, last antennomere filiform, 3.75–4.25 times as long as wide (Fig. 38), its ventral side smooth. Pronotum trapezoidal, ca. 1.81–2.03 times as wide as long. Sides almost straight, in some specimens with indistinct obtuse angle behind the middle (Fig. 58). Pronotum with two oval shallow depressions on the disc, and indistinct depressions medially also along anterior and posterior margin. Punctation consists of sparse moderately coarse setigerous punctures on the disc, which become slightly coarser and denser laterally. Elytra with numerous longitudinal rows of moderately large punctures, and very fine interstitial setigerous punctures. Ventral part with fine punctures, which are sparse medially and become larger and denser laterally. Last

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abdominal ventrite laterally with indistinct sinuation before apex. Male. Aedeagus with phallobase of the same length as parameres. Parameres simple, broad, their subbasal hook is missing, but the subapical hook is developed and obtuse. Median lobe narrowly lanceolate (Fig. 78). Female. Similar to male in habitus, but larger. Antenna shorter, and more slender. Distribution. The species is known from upland areas of Nepal and northern India (Uttaranchal, Sikkim, West Bengal). The occurrence in Bhutan needs to be verified by precisely localised specimens.

Eulichas sundaensis Hájek, sp. nov. (Figs. 18–19, 39, 59, 79) Type locality. Indonesia, Sumatra Isl. Lampung prov., Bukit Barisan Selatan Nat. Park, 5 km SW of Liwa, ca. 05°04’S 104°04’E, 600 m. Type material. 6 specimens — Holotype ♂ (NMPC), labelled: “S SUMATRA: Lampung prov. / BUKIT BARISAN SELATAN N.P.; / 5°4’S 104°4’E; ±600m;5km SW / Liwa; J. Bezd ě k leg.; 7.–17.ii.2000 [printed]”. Paratypes: 1♂ (no. 1), same label data as holotype (NMPC); 1♂ (no. 2), “SOUTH SUMATRA / Lampung Prov. / Wonosoko, Sdajo / lgt. P. Moric 12.iii.2000 [printed]” (NMPC); 1♂ (no. 3), “W. Sumatera, 500–800 m / HARAU VALLEY env. / 6–7.2005, loc coll [printed]” (NMPC); 1♀ (no. 4), “W. SUMATERA / Mt. Sanggul, 1200 m / Landai, 6 – 7.2004 [printed]” (NMPC); 1♀ (no. 5), “West Sumatera / MT. TALANG / 3–4.2005, 1000–1500m [printed]” (NMPC). Additional material studied. 4 specimens — INDONESIA: JAVA: 1♂ (MNHN); 1♂ 1♀, West Java, Tjikurai Mt., 4000’, 1892, H. Fruhstorfer (MNHN, NMPC); 1 ♀ , West Java, Gunung Papandayan Mt. (RMNH). Description. Habitus elongate, fusiform. Body colouring brownish-red in old speciemens from Java, and from brown to brown-blackish in recently collected material. Pale part of setation consists of recumbent yellowish setae covering uniformly pronotum, and forming indistinct ocellations on elytra (Fig. 18–19). Measurements. Males: 18–23 mm (holotype 23 mm); females: 22–27 mm. Head punctation consists of sparse, irregularly distributed moderately large setigerous punctures. Antenna slender, last antennomere filiform, 4.57–6.86 times as long as wide (Fig. 39), its ventral side smooth. Pronotum trapezoidal, ca. 1.73–1.87 times as wide as long. Sides regularly rounded in anterior half, then very slightly incised; hind angles distinctly prominent (Fig. 59). Pronotum with two indistinct rounded depressions on the disc. Punctation consists of sparse moderately coarse setigerous punctures on the disc, which become slightly coarser and denser laterally. Elytra with numerous longitudinal rows of moderately large punctures, and very fine interstitial setigerous punctures. Ventral part with fine punctures, which are sparse medially and become larger and denser laterally. Last abdominal ventrite laterally regularly rounded to apex. Male. Aedeagus with phallobase shorter than parameres. Parameres simple, relatively slender, subparallel. Their apical part (between subapical hook and apex) is relatively short. Subbasal parameral hook is missing, but the subapical hook is developed, slightly prominent. Median lobe narrowly lanceolate (Fig. 79). Female. Similar to male in habitus, but larger. Antenna shorter, and more slender. Remarks. Four old specimens from Java, which represent the only known Eulichas from Java so far, differ from the typical specimens in slightly smaller body length, brownish-red colouration of cuticle, and almost uniform body setation without any ocellations. As I was not able to resolve whether this is only intraspecific variability, I did not designate them as paratypes. Differential diagnosis. E. sundaensis sp. nov. is not similar to any other species of the E. sikkimensis species complex. It could be easily distinguished based on rather long pronotum with sides almost rounded anteriorly; slender antenna with long filiform last antennomere; slender, parallel sided parameres with short



35

apical part; and slender, narrowly lanceolate median lobe. Collection circumstances. Specimens from southern Sumatra were collected at light (J. Bezděk pers. comm., 2000), while specimens from western Sumatra were swept from vegetation at night (S. Jákl pers. comm., 2008). Distribution. A species known from several localities in western and southern Sumatra, and western Java. Etymology. The name is derived from “Sunda” Islands, where the new species occurs.

Eulichas wewalkai Hájek, sp. nov. (Figs. 20, 40, 60, 80) Type locality. Nepal, 30 km NW of Pokhara, Birethanti, 1100 m. Type material. Holotype ♂ (NHMW), labelled: “Nepal, 30 km NW Pokhara / Birethanti 1100 m / leg. Wewalka 4.5.1984 (N3) [printed]”. Description. Habitus elongate, fusiform. Body colouring brown. Pale part of setation consists of recumbent yellowish-grey setae forming very indistinct ocellations on pronotum and elytra (Fig. 20). Measurements. Male: 14 mm. Head punctation consists of sparse, irregularly distributed moderately large setigerous punctures. Antenna long, slender, last antennomere slightly club shaped, 2.63 times as long as wide (Fig. 40), its ventral side smooth. Pronotum trapezoidal, ca. 1.76 times as wide as long. Sides almost straight in anterior half, with obtuse angle behind the middle, and prominent hind angles (Fig. 60). Pronotum with two rounded shallow depressions on the disc, and an indistinct depression also medio-basally. Punctation consists of sparse moderately coarse setigerous punctures on the disc, which become slightly coarser and denser laterally. Elytra with numerous longitudinal rows of moderately large punctures, and very fine interstitial setigerous punctures. Ventral part with fine punctures, which are sparse medially and become larger and denser laterally. Last abdominal ventrite laterally with indistinct sinuation before apex. Male. Aedeagus with phallobase of the same length as parameres. Parameres simple, slender, somewhat constricted near the middle. Their subbasal hook is missing, but the subapical hook is well developed. Median lobe broadly lanceolate (Fig. 80). Female. Unknown. Differential diagnosis. E. wewalkai sp. nov., although of smaller size, is very similar to E. sikkimensis and E. uniformis in habitus. It can be distinguished from E. uniformis based on absence of a parameral subbasal hook, and from E. sikkimensis based on shorter last antennomere, slender parameres constricted near the middle, and broadly lanecolate median lobe. Distribution. So far known only from the type locality in mountain area of central Nepal. Etymology. The new species is dedicated to its collector, my colleague Günther Wewalka (Vienna, Austria), a specialist on Dytiscidae.

The Eulichas fasciolata species complex The complex contains four species from the Greater Sunda Island of Borneo. They differ from all other Eulichas species in the relatively short parallel sided parameres with a characteristic twisted apex. The subapical parameral hook is well developed, the subbasal hook is reduced. The median lobe is narrowly lanceolate, always longer than the parameres. The phallobase is about the same length as parameres and median lobe combined. Eulichas fasciolata (Fairmaire, 1898)

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(Figs. 21, 41, 61, 81) Lichas fasciolata Fairmaire, 1898: 388 (original description, Kina-Balu). Eulichas fasciolata (Fairmaire, 1898): Pic 1914: 11 (catalogue); Jäch 1995: 363 (catalogue).

Type locality. “Kina-Balu” [Sabah state, Malaysia]. Type material. Not found in MNHN. Additional material studied. 121 specimens — MALAYSIA: SABAH: 1♀ (SMTD); 2♀♀, coll. C. Felsche (SMTD); 26♂♂ 13♀♀, Kinabalu (DEIC, HNHM, MNHN, NHMB, SMTD); 6 ♂♂ 8♀♀ , Mount Kinabalu, v.–viii.1903, J. Waterstradt (MNHN, RMNH); 2♂♂ , Kinabalu National Park, Poring, 500 m, 29.xi.–2.xii.1996, W. Schawaller leg. (SMNS); 4♂♂ 2♀♀, Crocker Range [Mts.], iv.1990 (MCSN); 4♂♂, Crocker Range Mts., 50 km E Kota Kinabalu, Gunung Emas [Mt.], 16.–27.iv.1993, I. Jeniš leg. (IJCN, NHMW, NMPC); 1♂ , Keningau, 800 m, iv.1987 (NMPC); 6♂♂ 1♀ , Crocker Range, Keningau, v.1993 (NMPC); 1♂, 8,5 miles NW of Keningau, Kimanis road, at light, 11.v.1982, D. Burckhardt leg. (MNHG); 3♂♂, Keningau – Kimanis, km 25, iv.1994 (NHMW); 1♂, NW of Keningau, C.R.Park, Kimanis road, 10–15 km oberh., Park HQ, 05°26’N 116°05’E, 1100–1300 m, 2.–4.ix.1998, D. Bratsch & C. Häuser leg. (SMNS); 4♂♂ 3♀♀, Crocker Range, 16 miles NW of Keningau, 8.ix.1982, S. Nagai leg. (EUMC); 1♂, same label data, but 12.ii.1982 (EUMC); 2 ♂♂, same label data, but 1400 m, 2.–26.iv. 1984 (EUMC); 1♀, same label data, but 4.iv.1984 (EUMC); 1 ♂ , same label data, but 9.iv.1984 (EUMC); 1 ♂ , same label data, but 20.iv.1984 (EUMC); 1♂, Bunsit, 13.iii.1983, S. Nagai leg. (EUMC); 2♂♂, 105 km S Beaufort, Long Pasia Area, airstrip Long Pasia, 04°24’N, 115°43’E, 1000 m, at light, Recultivated area, 16.iv.1987, J. van Tol & J. Huisman leg. (RMNH); 3♂♂, 70 km W of Lahad Datu, Danum valley, 150 m, at light, 13.xii.1989, M.J. & J.P. Duffels leg. (ZMAN); 1 ♂ , Batu Punggul [ca. 04°38’N 116°35’E], Banjaran Maitland, 25.–27.v.1995, I. Jeniš leg. (MNHG); 1♂ , 50 km S Tomani, 4.v.1999, M. Snížek leg. (VKCZ); 1♀ , Trus Madi, 2200 m, 22.iv.1990, Martini leg. (NMPC); 2♂♂ 1♀, Mount Trus Madi, 1100 m, iv.2000, K. & B. Martini leg. (VKCZ); 6♂♂, Mt. Trus Madi, v.2004, S. Chew leg. (NMPC); 1♂, same data, but 3.iii.2005 (BMNH); 2♂♂, Mamut, 17.v.1979, N. Nishikawa leg. (EUMC); 1♂ , Sepilok, Sandakan, 6.ix.1982, S. Nagai leg. (EUMC); 1♂ , 80 km E of Ranau, Telupid, 800 m, 20.iv.1979, S. Nagai leg. (EUMC); 1♂, Tinamantawaran, near Ranau, 16.iv.1983, S. Nagai leg. (EUMC). SARAWAK: 1♂, Gunong Mulu National Park, 150 m, at light, ii.1978, J.D. Holloway et al. (BMNH); 2♀♀ , same label data, but 4 th Division, 150–200 m, v.–viii.1978, P.M. Hammond & J.E. Marshall leg. (BMNH). Description. Habitus elongate, fusiform. Body colouring redish-brown to brown. Pale part of setation consists of recumbent brownish-yellow setae covering uniformly head, pronotum and ventral part, and forming rather indistinct ocellations on elytra (Fig. 21). Measurements. Males: 23–32 mm; females: 30–35 mm. Head punctation consists of sparse moderately large setigerous punctures. Antenna long, slender, last antennomere narrowly oval, 3.19–3.58 times as long as wide (Fig. 41), its ventral side smooth. Pronotum trapezoidal, ca. 1.90–1.91 times as wide as long. Sides almost straight, in some specimens with indistinct sinuation behind the middle, hind angles prominent (Fig. 61). The disc convex with two lateral and one basal shallow depression. Punctation consists of sparse moderately coarse setigerous punctures on the disc, which become slightly coarser and denser laterally. Elytra with numerous longitudinal rows of moderately large punctures, and fine interstitial setigerous punctures. Ventral part with fine punctures, which are sparse medially and become larger and denser laterally. Last abdominal ventrite laterally regularly rounded to apex. Male. Aedeagus with phallobase shorter than parameres. Parameres parallel sided, with twisted emarginated apex; their distal part rather short. Median lobe lanceolate, with thin apical part; exceeding parameres (Fig. 81). Female. Similar to male in habitus, but larger. Antenna shorter, and more slender.



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Remarks. Although the type material of the species was not found/recognised in the MNHN, the large number of additional specimens from Kinabalu deposited in various collections has allowed confident identification and redescription of the species. Collection circumstances. Collected at light in the primary forest. Distribution. Known from the Malay part of the Greater Sunda Island of Borneo – Sabah and Sarawak.

Eulichas jakli Hájek, sp. nov. (Figs. 22, 42, 62, 82) Type locality. Indonesia, southern Kalimantan, Kandangan district, 17 km NE of Loksado. Type material. Holotype ♂ (NMPC), labelled: “INDONESIA / S. Kalimantan / Kandangan district // 17 km NE Loksado / 15.11.[19]97/15.1.[19]98 / St. Jákl lgt [printed]”. Description. Habitus elongate, fusiform. Body colouring brown. Pale part of setation consists of recumbent brownish-yellow setae covering uniformly head, pronotum and ventral part, and forming typical ocellations on elytra (Fig. 22). Measurements. Male: 21 mm. Head punctation consists of sparse moderately large setigerous punctures. Antenna long, slender, last antennomere filiform with apical elongation, 4.71 times as long as wide (Fig. 42), its ventral side smooth. Pronotum trapezoidal, ca. 1.95 times as wide as long. Sides almost straight with indistinct obtuse angle behind the middle, hind angles prominent (Fig. 62). The disc convex. Punctation consists of sparse moderately coarse setigerous punctures on the disc, which become slightly coarser and denser laterally. Elytra with numerous longitudinal rows of moderately large punctures, and fine interstitial setigerous punctures. Ventral part with fine punctures, which are sparse medially and become larger and denser laterally. Last abdominal ventrite laterally regularly rounded to apex. Male. Aedeagus with phallobase of the same length as parameres. Parameres broad, with twisted apex; their distal part relatively long. Median lobe narrowly lanceolate, exceeding parameres (Fig. 82). Female. Unknown. Differential diagnosis. E. jakli sp. nov. could be easily distinguished from all other species of the E. fasciolata complex based on small body length, the very thin and long last antennomere, and characteristic shape of aedeagus with long distal part (between apex and subapical hook) of parameres. Distribution. So far known only from the type locality in southern Kalimantan, Indonesia. Etymology. The new species is dedicated to its collector, my friend Stanislav Jákl (Praha, Czech Republic), a specialist on Cetoniidae.

Eulichas subocellata (Fairmaire, 1898) (Figs. 23, 43, 63, 83) Lichas subocellata Fairmaire, 1898: 388 (original description, Kina-Balu). Eulichas subocellata (Fairmaire, 1898): Pic 1914: 11 (catalogue); Jäch 1995: 363 (catalogue).

Type locality. “Kina-Balu” [Sabah state, Malaysia]. Type material. Lectotype ♀ (MNHN), by present designation: “Kina-Balu / Bornéo. [printed] // TYPE [red label, printed] // M USÉUM P ARIS / 1952 / C OLL . R. O BERTHUR [yellow label, printed] // Lichas / subocellatus / Fairm. [handwritten] // LECTOTYPUS / LICHAS / subocellata Fairmaire, 1898 / Jiří Hájek des. 2002 [red label, printed] // EULICHAS (s. str.) / subocellata (Fairmaire, 1898) / Jiří Hájek det. 2002 [printed]”. Paralectotypes: 1♀ , same label data as lectotype; 2♀♀ : “Kina Balu. / N. Borneo. [printed] //

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MUSÉUM PARIS / 1952 / COLL. R. OBERTHUR [yellow label, printed] // SYNTYPE [red label, printed]”, and the respective paralectotype label (MNHN). Number of syntypes unknown. I designate a lectotype to fix the identity of this species, as available taxonomic works do not allow an unambiguous identification of specimens. Additional material studied. 113 specimens — MALAYSIA: SABAH: 7♂♂ 7♀♀, Kinabalu (DEIC, HNHM, IRSNB, MNHN, SMTD, ZMHB); 2♀♀, same label data, but 1500 m, H. Rolle (DEIC, HNHM); 2♂♂ 4♀♀, Mount Kinabalu [ca. 06°04’N, 116°36’E] (BMNH); 8♂♂ 8♀♀, same label data, but v.–viii.1903, J. Waterstradt (BMNH, MNHN, NHMB, RMNH); 1♂, Gunung Kinabalu, Sg. Liwagu nr Kundasang bridge, 06°00’N, 116°34’E, 1185 m, 23.xi.1986, J. Huisman leg. (RMNH); 7♂♂ 5♀♀, Kinabalu NP, Sayap, 1000 m, 25.–29.XI.1996, W. Schawaller leg. (NMPC, SMNS); 1♂, Kinabalu Nat. Park, Sayap, 6°10’N 116°34’E, primary forest edge, 950 m, at light, 8.iii.2001, J.P. Duffels & M.A. Schouten leg. (ZMAN); 3 ♂♂ , Tinamantawaran, near Poring, Ranau, i.1983; 1♀, S of Tambunan, 650 m, 4.–5. & 13.vii.1991, J. Haxaire leg. (NHMW); 1♂, Crocker Range Mts., E. Khoo leg. (VKCZ); 1♀, Crock[er] Range, 17.ii.1989 (MCSN); 1♂, same label data, but iv.1990 (EUMC); 1♂, same label data, but 5.v.2005, S. Chew leg. (BMNH); 1♂, Crocker Range, 16 Mail [miles?] NW of Keningau, 1400 m, 20.iv.1984, S. Nagai leg. (EUMC); 1♂, same label data, but 2.–13.v.1984 (EUMC); 18♂♂ 1♀, Crocker Range Mts., 50 km E Kota Kinabalu, Gunung Emas [Mt.], 16.–27.iv.1993, I. Jeniš leg. (IJCN, NHMW); 1♂, Crocker Mt., Gunong Emas, 6.–21.v.1995, I. Jeniš leg. (NMPC); 1♀, same data, but J. Stolarczyk leg. (NMPC); 1♂, Kundasang, 1500 m, 4.–8.ix.1994, C.L. Li leg. (NHMW); 12♂♂, Banjaran Crocker Mts., 16 km SW Gunung Alab, 790–850 m, 4.–9.v.1996, M. Štrba & R. Hergovits leg. (NHMW, NMPC); 1♀, Mt. Trus Madi, v.2004, S. Chew leg. (NMPC); 5♂♂ 2♀♀, same data, but 3.iii.2005 (BMNH, NMPC); 1♂, Tawau hill, along Sungai Tawau near trail to Air Panas, 300 m, at light, 28.iii.2001, J.P. & M.J. Duffels leg. (ZMAN); 1♂, Tawau, 20.iii.2006 (NMPC). SARAWAK: 3♂♂, Mt. Dulit, moss forest, 22.x.1932, Oxford University Expedition: B.M. Hobby & A.W. Moore (BMNH); 6♂♂ , 4th Division, Gunung Mulu National Park, at light, 150–200 m, v.–viii.1978, P.M. Hammond & J.E. Marshall leg. (BMNH); 3♂♂ 1♀, 105 km S Beuafort, Long Pasia Area, Sg. Maga nr confl. Sg. Pasia, 04°26’N, 115°40’E, 1210 m, at light, along larger fast running stream in lower montane evergreen rain forest, 2.–7.iv.1987, J. van Tol & J. Huisman leg. (RMNH). Description. Habitus elongate, fusiform. Body colouring dark brown. Pale part of setation consists of recumbent grey-whitish setae forming indistinct ocellation on head, pronotum and ventral part, and characteristic distinct ocellations on elytra (Fig. 23). Measurements. Males: 24–26 mm; females: 31–34 mm (lectotype 33 mm). Head punctation consists of sparse moderately large setigerous punctures. Antenna long, slender. Last antennomere narrowly oval with apical elongation, 3.40–4.46 times as long as wide (Fig. 43), its ventral side smooth. Pronotum trapezoidal, ca. 1.95–2.09 times as wide as long. Sides slightly, almost regularly rounded (Fig. 63). The disc convex, in some specimens with irregular shallow depressions. Punctation consists of sparse moderately coarse setigerous punctures on the disc, which become slightly coarser and denser laterally. Elytra with numerous longitudinal rows of moderately large punctures, and fine interstitial setigerous punctures. Ventral part with fine punctures, which are sparse medially and become larger and denser laterally. Last abdominal ventrite laterally regularly rounded to apex. Male. Aedeagus with phallobase shorter than parameres. Parameres parallel sided, with twisted emarginated apex; their distal part rather short. Median lobe narrowly lanceolate, with thin apical part; exceeding parameres (Fig. 83). Female. Similar to male in habitus, but larger. Antenna shorter, and more slender. Collection circumstances. Collected at light in the primary forest. Distribution. Known from the Malay part of the Greater Sunda Island of Borneo – Sabah and Sarawak. Eulichas villosa Hájek, sp. nov. REVISION OF EULICHAS II


39

(Figs. 24, 44, 64, 84) Type locality. Malaysia, Sabah, Gunung Emas Mt., 1700 m. Type material. 18 specimens — Holotype ♂ (NMPC), labelled: “Malaysia-Borneo / Sabah 21.3.– 20.4.1996 / GUNUNG EMAS 1700m / lgt. J. Kadlec [printed]”. Paratypes: 1♂ (no. 1) 2♀♀ (nos. 2–3), same label data as holotype (NMPC); 1♀ (no. 4), “Borneo 15/27.4.1993 / Sabah Crocker Mt. / Gunong Emas env. / Jenis & Strba leg [printed]” (NHMW); 2♂♂ (nos. 5–6), “BORNEO-SABAH / Crocker Mt. 500–1900m / Gunong Emas / 6.–21.5.1995 / Ivo Jeniš leg. [printed]” (MHNG, NMPC); 1♀ (no. 7), “GUNUNG EMAS / 7.2.[19]99 / BORNEO / LEG.KONDLER [printed]” (NMPC); 1♂ (no. 8), BORNEO / GUNUNG ALAB / 19. – 24. 4. 2006 / B.MAKOVSKY Lgt. [printed]” (NMPC); 1♂ (no. 9), “(Malaysia) / headquarters / (alt.1500– 1700m) / Mt. Kinabalu / Sabah / III/27.1976 / S.NAGAI leg. [printed]” (EUMC); 1♀ (no. 10), samel label data, but “IV-15.1976” (NMPC); 1♀ (no. 11), “(EAST MALAYSIA) / Park Headquarters / near Mt. Kinabalu / Sabah, (alt. 1700m) [printed] / III - 25. [handwritten] 1979 / SHINJI NAGAI leg. [printed]” (EUMC); 1♂ (no. 12), “Headquarters / Borneo Sabah / April 18, 1979 / N. Nishikawa [printed] // Eulichas / subocellata / (Fairmaire) [handwritten] / DET. M. SATO 1980 [printed]” (EUMC); 1♀ (no. 13), same label data, but “April 25” (EUMC); 1♀ (no. 14), “(N BORNEO) / Headquarter / Mt. Kinabalu / 3. V. 1980 / M. & A. Sakai [printed]” (EUMC); 1♀ (no. 15), same label data, but “4. V. 1980” (EUMC); 1♂ (no. 16), same label data, but “6. V. 1980” (NMPC); 1 ♀ (no. 17), “BORNEO SABAH Mt. / Kinabalu Nat.Pk. / HQ 1560 m 15–24. / V.[19]87 A.Smetana [printed]” (MHNG). Description. Habitus elongate, fusiform. Body colouring brown. Pale part of setation consists of recumbent yellowish or greyish setae covering uniformly head, pronotum and ventral part, and forming rather indistinct ocellations on elytra (Fig. 24). Measurements. Male: 24–27 mm (holotype 26 mm); females: 29–33 mm. Head punctation consists of sparse moderately large setigerous punctures. Antenna long, slender, last antennomere almost filiform, 3.80–4.44 times as long as wide (Fig. 44), its ventral side smooth. Pronotum trapezoidal, ca. 1.87–2.00 times as wide as long. Sides almost straight, with indistinct sinuation behind the middle, hind angles prominent (Fig. 64). The disc convex with two lateral and one basal shallow depression. Punctation consists of sparse moderately coarse setigerous punctures on the disc, which become slightly coarser and denser laterally. Elytra with numerous longitudinal rows of moderately large punctures, and fine interstitial setigerous punctures. Ventral part with fine punctures, which are sparse medially and become larger and denser laterally. Last abdominal ventrite laterally regularly rounded to apex. Male. Aedeagus with phallobase shorter than parameres. Parameres slightly narrowed in the middle, with twisted emarginated apex. Median lobe lanceolate, slightly constricted before apex; exceeding parameres (Fig. 84). Female. Similar to male in habitus, but larger. Antenna shorter, and more slender. Differential diagnosis. The new species is very similar to E. fasciolata. However, E. villosa sp. nov. is sligthly smaller and more slender, with denser body setation; antennomeres are slender and more elongate. In addition, E. villosa sp. nov. differs in the shape of aedeagus: parameres are slightly narrowed medially, their subapical hook is less prominent; median lobe is more slender, slightly constricted before apex. Distribution. So far known only from several mountain localities in Sabah, Malaysia. Etymology. The Latin word for “villose” refers to peculiar dense body setation of the species.

Species not revised

40


HÁJEK

Eulichas trapezicollis (Fairmaire, 1891) Lichas trapezicollis Fairmaire, 1891: CXXIX (original description, Darchiling). Eulichas trapezicollis (Fairmaire, 1891): Pic 1914: 11 (catalogue); Jäch 1995: 363 (catalogue).

Type locality. “Darchiling” [Darjeeling district, West Bengal, India]. Remarks. Eulichas trapezicollis is almost certainly a senior synonym of E. sikkimensis or E. uniformis, which both occur in Darjeeling. Both species are very similar in habitus, and could be reliably distinguished only based on shape of last antennomere or male genitalia. Unfortunately the original description did not mention any of these diagnostic characters, and the type material of E. trapezicollis seems to be lost. Therefore I am not able to establish the formal synonymy. Type specimens should be deposited in MNHN, but I failed to find them there during my visits. However, I still believe, that more searching could reveal them, and therefore do not designate a neotype, but leave the status of E. trapezicollis unresolved.

Discussion The present revision raises the number of known Eulichas species to 42. The identity of one species – E. trapezicollis remains doubtful. It confirms the preliminary division of Eulichas into two species groups as suggested by Jäch (1995). The E. dudgeoni species group displays several groundplan features of the family (apical antennomere not widened; aedeagus slender, phallobase not prolonged, parameres about the same length, or slightly shorter than median lobe). However, absence of any significant apomorphy does not allow the monophyly of the group to be confirmed. As in the first part of my revision (Hájek 2007), I added the species complex criteria for easier orientation within the genus. In contrast to the E. funebris species group, the monophyly of the species complexes remains uncertain, with the exception of the E. fasciolata species complex, which is well characterized by a single apomorphy: the twisted apex of the parameres. Partly, the species of the E. dudgeoni species group show even more variability than those of the E. funebris species group. As in the first part of the revision (Hájek 2007), I am using, more or less, a combination of three characters for taxonomic purposes: (1) shape of the last antennomere; (2) shape of pronotum; and (3) shape of the aedeagus. These characters are in some cases augmented with the general habitus of species or their body setation. However, any of the characters mentioned above can vary in some specimens, and therefore identification of single specimens or females without males is often impossible. The problematic variability is also a reason why a large proportion of the specimens examined are not included in the type material of the newly described species.

Zoogeographical considerations Although our information about Eulichas is still fragmentary and incomplete, large territories within its area of distribution are still unexplored, and a lot of collected material is not precisely localised, several basic trends of Eulichas distribution can be outlined here. The presently known distribution of Eulichas covers the area from Himachal Pradesh in north-western India, Nepal, Bhutan and north-eastern India, central and southern China, the whole territory of continental south-eastern Asia (so far except for Cambodia), the Greater Sunda Island Sumatra (and the surrounding islands Belitung, Nias, Siberut), Java and Borneo, and most of the islands of the Philippines. Within the whole area of distribution, Eulichas species occur predominantly in mountain areas, the highest diversity is in Eastern Himalayas (Nepal, Bhutan, India), Annam Highlands (Thailand, Laos, Vietnam), Cameron Highlands (Malaysia), Barisan Mts. (Sumatra) and Crocker Range (Borneo). All of the currently defined Eulichas species complexes, except for the E. sikkimensis complex are REVISION OF EULICHAS II


41

characteristic for any well defined region within the area of distribution of Eulichas. These regions (summarised in Table 1) are as follows: TABLE 1. Type of distribution of Eulichas species. Type of distribution Border area between Palaearctic and Oriental zoogeographical regions E. funebris complex (5 sp.) E. dudgeoni complex (7 sp.) E. sikkimensis complex (3 sp.)

Eastern Himalayas

Central and south-eastern China

species

n

%

E. gigantea E. meghalayensis E. sikkimensis E. uniformis E. wewalkai

5

12

4

10

6

14.5

10

24

1

2.5

6

14.5

E. bertiae E. dudgeoni E. funebris E. tenuicornis

Northern parts of Thailand, Laos and Vietnam, E. alesbezdeki and Chinese Yunnan prov. E. jendeki E. siamensis E. similis E. tonkinensis E. undulata Continental SE Asia – Annam Highlands E. milleri complex (2 sp.) E. pacholatkoi complex (4 sp.) E. sikkimensis complex (1 sp.) sg. Forficulichas (3 sp.)

Malay Peninsula (+ Sumatra) E. jaechi complex (4 sp.) E. sikkimensis complex (4 sp.)

E. dembickyi E. haucki E. horaki E. janbezdeki E. kubani E. milleri E. oborili E. pacholatkoi E. pantherina E. phoca

Tenasserim

E. birmanica

Cameron Highlands

E. incisicollis E. jaechi E. strbai E. tanahrata E. robusta E. serricornis E. sausai

1

2.5

Sumatra and Java E. mediocris complex (2 sp.) E. sikkimensis complex (1 sp.)

Malaysia + Sumatra

E. mediocris E. minuta E. sundaensis

3

7.5

Borneo E. fasciolata complex (4 sp.)

E. fasciolata E. jakli E. subocellata E. villosa

4

10

N Borneo + Philippines E. sikkimensis complex (1 sp.)

E. baeri

1

2.5

1. Border area between Palaearctic and Oriental zoogeographical regions. The transitional zoogeographical area covers the territory of northern India, Nepal, China and northern parts of Thailand, Laos and Vietnam. It is very characteristic in Eulichas fauna composition. Two species complexes: E. funebris and E. dudgeoni occur only in that region. The area could be further split to three smaller regions: i) Eastern Himalayas; ii) central and south-eastern China; and iii) the northernmost part of Annam Highlands. The last mentioned region continues in following area. 2. Continental south-eastern Asia – Annam Highlands. This area represents the current center of diversity

42


HÁJEK

of Eulichas. Altogether 10 species from both subgenera and both species group (if counted together with the northern most part of Annam Highlands, then 16 species – more than one third of described taxa) occur there. The characteristic species complexes are those of E. milleri and E. pacholatkoi, and also subgenus Forficulichas. 3. Malay Peninsula. The area with highest diversity in Cameron Highlands. The E. jaechi species complex is characteristic of this area. One species of the E. sikkimensis complex – E. sausai sp. nov. – represents the connection between Malay Peninsula and northern Sumatra. 4. The Sunda Islands Sumatra and Java, and surrounding small islands. A relatively less speciose area. It is characterized by the E. mediocris species complex. The uncertain position of Java, with only a single known species, could be due to the high rate of deforestation and destruction of habitats. 5. The Greater Sunda Island of Borneo is characterized by the occurrence of four species of the E. fasciolata species complex. 6. Philippines. The Philippines, with a single species of the E. sikkimensis species complex – E. baeri – has a specific position within Eulichas. The species is known from most of Philippine Islands and also from the northern part of Borneo. However, the origin of the species and its relationships with other species are unknown.

Acknowledgements I am obliged to all colleagues mentioned in the list of collections for putting at my disposal specimens from their collections, or loaning the material in their care. I also wish to thank to all my friends and colleagues for donation of their Eulichas material collected during various expeditions to south-eastern Asia – this revision would not have been possible without their gratuitous support. I am indebted to Pavel Sláma (Nové Hrady, Czech Republic) for his excellent drawings. I am very grateful to Maxwell V. L. Barclay (BMNH) and Manfred A. Jäch (NHMW) for their help and valuable comments to the manuscript. The present study was partly supported by the Ministry of Culture of the Czech Republic (MK00002327201) and by SYNTHESYS project (AT-TAF 1087). Work on Eulichas from BMNH was supported by the NHM Special Funds Grant “Increasing access to the Elateriformia collections”.

References Fairmaire, L. (1891) Descriptions de Coléoptères des Montagnes de Kashmir (Suite). Bulletin ou Comptes-Rendus des Séances de la Société Entomologique de Belgique, 1891, CXXI–CXXXIV. Fairmaire, L. (1898) Descriptions de Coléopterères d’Asie et Malaisie. Annales de la Société Entomologique de France, 67, 382–400. Hájek, J. (2007) Revision of the genus Eulichas Jacobson, 1913 (Coleoptera: Eulichadidae) I. Introduction, morphology of adults, key to subgenera and species groups, and taxonomy of E. funebris species group. Zootaxa, 1260, 1–35. Hájek, J. (2008) A new species of Eulichas (Coleoptera: Eulichadidae) from Laos. Acta Entomologica Musei Nationalis Pragae, 48, 67–72. Ivie, M.A. & Jäch, M.A. (2002) A new species of Eulichas Jacobson from Vietnam (Coleoptera: Eulichadidae). Koleopterologische Rundschau, 72, 165–168. Jäch, M.A. (1995) Eulichalidae: Synopsis of the species of the genus Eulichas Jacobson from China, Laos and Vietnam (Coleoptera), pp. 359–388. In: Jäch, M.A. & Ji, L. (Eds.) Water Beetles of China. Vol. 1. Wien: Zoologischbotanische Gesellschaft in Österreich and Wiener Coleopterologenverein, 410 pp. Jäch, M.A. & Hájek, J. (2006) Eulichadidae, p. 455. In: Löbl, I. & Smetana, A. (Eds.) Catalogue of Palaearctic Coleoptera, Volume 3, Scarabaeoidea - Scirtoidea - Dascilloidea - Buprestoidea - Byrrhoidea. Stenstrup: Apollo Books, 690 pp. Pic, M. (1911) Coléopterès exotiques nouveaux ou peu connus. L’Echange, Revue Linnéenne, 27, 142–144. Pic, M. (1913) Coléoptères exotiques en partie nouveaux. L’Echange, Revue Linnéenne, 29, 106–110. Pic, M. (1914) Pars 58: Dascillidae, Helodidae, Eucinetidae. In: Schenkling, S. (Ed.) Coleopterorum Catalogus, Berlin:



43

W. Junk, 65 pp. Pic, M. (1915) Descriptions abrégées diverses. Mélanges Exotico-Entomologiques, 12, 3–20. Pic, M. (1924) Nouveautés diverses. Mélanges Exotico-Entomologiques, 42, 1–32. Pic, M. (1933) Nouvealtés diverses. Mélanges Exotico-Entomologiques, 62, 1–36.

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