Dec 31, 1982 - River in. Venezuela. Morphometric and meristic data of several ...... Cojedes,. Rio. Salinas, a tributary of Rio. Pao Viejo,. N.E. of. El. Baul.
Beaufortia INSTITUTE
OF TAXONOMIC ZOOLOGY
UNIVERSITY
Vol. 32
(ZOOLOGICAL MUSEUM)
OF AMSTERDAM
8
no.
Dentectus mailed
a
barbarmatus,
catfish
from
the
new
Siluriformes,
(Pisces,
genus
Orinoco
December
31,
species
of
and of
Basin
1982
Venezuela
Loricariidae)
F.J. Martin Salazar de Estudios
Departamento
and
Venezuela,
Universidad Simon Bolivar,
Ambientales,
Laboratorio
de
de
Instituto
Ictiologia,
I.J.H. Isbrücker Taxonomische
voor
P.O. Box
& H.
Tropical,
de
Nijssen
Universiteit
Zodlogie,
van
Amsterdam,
The Netherlands
Amsterdam,
HC
1000
20125,
1081,
Universidad Central
Caracas
Venezuela,
Instituut
Zoologia
Caracas
Apartado 80659,
Abstract
Dentectus is
barbarmatus,
described
meristic with
from
a
data of several
Bleeker,
Planiloricaria
of
the
are
species
northern
are
of mailed catfish
discussed.
of
margin
presented It is
Rhadinoloricaria
1862,
Isbrücker,
and
genus
specimens
of the tribe
other genera
hemiodon
new
tributaries
and
the
of the
illustrations
assigned
Isbrücker
&
subfamily Loricariinae,
Orinoco are
River
given.
to the subtribe
Nijssen,
in
Venezuela.
The
tribe
of the
relationships
Planiloricariina, together Crossoloricaria
1974,
Loricariini, and
Morphometric new
genus
with Pseudo-
Isbrücker,
and
1979,
1971.
Resumen
Se
describe
Dentectus
Loricariini,
morfométricos discuten
las
hemiodon loricaria
barbarmatus
proveniente
de
merísticos
y
relaciones
Bleeker,
de
1862,
Isbrücker,
de
este
varios
de
en
la
género
la subtribu
y
especie norte
márgen
specímenes
nuevo
Rhadinoloricaria
1971,
género
un nuevo
tributarios
con
así
Isbrücker
&
late
Agustin
early
as
represented Ydpez. this Ten to
1968
in
a
the
was
years
on
of
collection for
the work,
when
initiated
Fernandez
different
they
Fernandez
the
the Loricariinae
interrupted during
later,
and
Yepez
publication
study
Unfortunately,
study
finish
in
misma
Nijssen,
a
de la
Orinoco,
ilustraciones
la
en
del
tribu;
subfamilia Venezuela.
holotipo
y
se
lo
y
tribu
Loricariinae, Se
incluyen
datos
algunos paratipos. junto
agrupa,
Crossoloricaria
1974,
Last Fernandez
senior author of the present as
Río
Isbrücker,
1979,
Se
Pseudo-
con
y
Plani-
Planiloricariina.
INTRODUCTION The
como
otros de
de Loricárido
del
reasons
long period.
were
Y£pez
preparing died.
study
year in
the
senior
collaboration
author with
restarted
Isbriicker
the
and
Nijssen. This is
mainly
the
with
Fernandez
general
of
a
species
very his
benefit
of
series of
He
accurate
specimens. gained
papers
dealing
Loricariinae in the
Yepez collection.
unpublished sketches
first of the
In
by
left
numerous
and
drawings addition
publishing
to
the
these
125
the
senior
author
with
gether
the
specimens,
his dez
Yepez
the
to-
necessary)
identification
of
to honour FernanSouth American
outstanding
and
Ichthyologist
still
and
also want
we
an
as
where data
of
pioneer
Ichthyological
Studies in Venezuela.
sorted out from
nating the
the
Both
Orinoco). one
another in
shape of
maxillae
River,
which
groups
were
slender
They
two
group
assigned
1854),
“Loricaria”
Together
“Loricaria”
distinct from
were
a
forthcoming
second as
it
with
(1904:
274,
nominate
in
variegata,
mina,
“L.”
1904,
the
forms
two
represent
a
different
group, as
defined
recognized
was
the
as
Dentectus
of
specimens
group "aff.
cf.
Loricaria
represents will be
of
a
species
described in
“L.”
laticeps,
Pseudohemiodon
included
IV,
Loricaria
pointed was
appeared, by
“L.” out
distinct
Regan in
B)
the
“Lori-
(viz.:
macromystax,
and
“L.”
la-
platycephala). especially
that
generic
level
at
Isbriicker,
and
and
In this
to
the
the
tion of Fernandez at
first
sight
specimens
after
this
These
structure of the of
the
lip,
the
the
ever,
new
with both
the
and
to have
few
to be
seems
upper
or
in
any
How-
related
closely
Pseudohemiodonand group
the
of
the Pseudo-
Loricariidae.
family
genus
base
the
of
the
as
armature
the
at
Planiloricaria,
the
very small
well
as
peculiar
and
all.
stained
genus of
any
is
collec-
teeth at
jaws and
barbels
in
of
genus
in the
no
filamentous barbels
hemiodon-group, other
had
both
pre-
considered which
are
characters
unknown in
are
the
Planiloricariina.
they proved
lips, and
maxillary
of
some
of
Loricariina,
cleared and
were
treatment
premaxillae.
new
1979, 1974a,
result
specimens
Yepez
minute teeth in
very
1975,
Nijssen, a
subtribe
apparently
Two
&
As
subtribe
those
paper,
some
1973,
Pseudohemiodon-group,
in
placed
assigned
now
1978).
the
present study,
1971,
Isbriicker
1981;
1976,
of
description
the
Planiloricaria.
genus
“Lori-
genus:
couplet
key,
was
1980, 1974b,
the
to
ACKNOWLEDGEMENTS
publications
species
the
(cf.
Thanks
to the kindness who
Leccia,
work
to
in
Zoologia de
his
laboratory of
Tropical
extensive
of
and
at
the
species
the
gave
under
his
been able to locate additional same
Francisco
Mago
Instituto
Universidad
who
collection
Dr
the senior author facilities
provided
Venezuela
Central to
access
care,
de
we
the
have
specimens of the
in the collection of Fernandez
as
Yepez. The senior author likes to thank the Decanato de
of
Investigaciones
Bolivar
several
subgenus
it
defined
paper.
caria”
Initially,
a
1758,
and
described
new
a
with
platycephala
to
group
herein
lamina”;
1968
Since
dealing
as
on
then
laticeps Regan,
seem
Pseudohemiodon which
of
pre-
species.
related
of Loricaria
labelled
L.
to
these
Linnaeus,
species,
The first
of
initially
the
barbels
two
1869.
Salinas
The
barbarmatus.
of
obvious
was
Gunther,
species,
representative
laticeps and
It
1868,
species
(1904).
new
of
general
neither the
on
easily identifiable
the other
a
One
Gunther,
these
Rio
and
by Regan as
with
in
into
in
as
viz.: “Loricaria”
caria” macromystax
collected
very
such
Loricaria
lamina
species
origi-
similar
subgenus Loricaria
the
by Regan (1904), (Kner,
flows
closely
were
of
species to
tributary
represented
groups
apparently of
(a
filamentous
lips.
both upper and lower that these
fishes
the dentaries and had remark-
on
and
ably long
senior
of
snout.
visible teeth
no
nor
River
head and
body,
had
groups
groups
appearance,
the
and
Yepez
two
Salinas
Portuguesa
and
viously
Fernandez
1968,
In
author
genera
spend
for
a
the
few
weeks
Taxonomische van
Universidad
in
Zoologie
support
the of
Simon
given
Instituut the
to
voor
Universiteit
Amsterdam.
The
drawings
sketched
by
completed were
the
financial
made
in
figs.
Agustin
by by
F. Mr
J. L.
I
and
Fernandez
Martin A.
van
2
were
Yepez
Salazar.
Figs.
der Laan
first and
3-4
(ZMA).
MATERIAL AND METHODS from
Loricaria
different
proposed,
126
taxonomic next
lato,
sensu
to
and
assignments
the
successively have
establishment
of
been new
Specimens Museo
de
are
deposited
Biologia,
in
collections
Universidad
of
Central
the de
Venezuela
Universiteit The
proposed
in
measurements
before
the
calculating
traction of
this
part
press)
elongation
which
is
it
various
enables
a
In
I
Diagnosis. cariinae,
included, Subfor
way
data
morphometric
some
those referred
above,
to
—
are
fol-
as
lows:
length/pelvic
I):
ratio of
in abdominal
in
fin
pelvic
spine,
40
in
expressed
length,—
abdominal scutelets
scutelets
spine (character
number of
(character 44):
posterior
anterior
+
transverse
series,—
of
SL,
snout
to
tip
of snout
ratios
pressed
least
pelvic
in
orbital
diameter
orbital width,
maxillary barbel, orbital
from
10): fin
pelvic
spine;
diam.
(character
orbital
diameter,
ex-
width,—
(character
ratio
37):
in
least
21):
(character
of
=
(character
ratios
width
body
25):
and
anterior
expressed at
width
in
anal at
orbital
dia-
posterior
HL,
(character
anal,
38):
expressed
in
of anal fin
7):
from
tip
the
from to
up
expanded
the
are
normally
the
lower
preserved
barbels
smooth.
are
maxillary.
up
are
present. from each
three teeth each,
to
the
in
ones
jaws
the
pre-
invisible
are
specimens:
specimens
In
small, elongate
widely separated
similar to
observed in
lip
enlarged
other
the
three
The teeth and
maxillary.
dermal
branchiostegal
crowns
distally incasing
they
which
in be
can
are
cleared
and stained.
Additional characters with
comparison
of the
Fin-ray
last
one
rays;
10, I
anal last
one
meaning
in
meaning
as
to its
fin
rays;
fin
only
dorsal
split
in
split
in
with
I, 5
nine
genera
1,6,
in
a
in
the
i
of
the
rays,
the
pectoral fin with I, 6 rays;
caudal fin with
specimens,
four
and
species.
other
base;
below
given
genera
known
fin
with
and
rays
are
related
counts
pelvic
of snout
spine; ratios expressed
and
other
with
Etymology.
preanal length (character
SL,—
each
of
small, widely separated
very
premaxillary
other,
I,
—
of
maxillary barbel)
Dentaries likewise
rictal
postanal length,—
to base
from
Loricariini:
excluding
postanal/body
for the
description
—
barbel
diameter
notches;
laterally,
Premaxillaries
each
and
inter-
of
margin
height
whereas
easily
anterior
excluding
expressed
36):
provided
the
margin
with
papillae
which teeth
—•
—
meter
ratio
SL,
diam.
notches,
posterior
(character
interorbital
interorbital/orbital of
tip
ratios
filaments of
entire
odontodes.
the
along
the
of
the
irregular
these barbels have
(except
&
dis-
long,
numerous
rather
membrane; usually
teeth
maximum in
from
9):
papilla;
anal
—
orb.
interorbital/max. of
of
to outer base of
expressed
ratio
(character
anus
center
in
expressed distance
to
to
snout
barbel
bearing
Barbels to
Isbriicker
following
lip with
almost
maxillary
reach
(3)
distance snout
the
barbels in several
and
lip
upper
Lori-
subfamily
(sensu
Externo-lateral base
rows.
(2)
179),
species
new
characters:
ossifications
table
genus
the
with
and lower
Upper
of
genus
Loricariini
1978:
tinguishing (i)
A
tribe
Nijssen,
the
abdominal
new
Dentectus barbarmatus
Type-species:
filamentous
table
39):
length.
Dentectus
from
comparison.
additional to
(character
length, expressed
SYSTEMATIC DESCRIPTIONS
have
ratios.
better
thoracic
spine
spine
ante-
ventrorostral
(in
this
fin
pectoral
180-
(1978:
(or
length/pectoral
of
and
those
as
produced,
a
al.
et
subtract
to
ratio in
same
Nijssen
snout
Isbriicker
extension)
&
thoracic
(ZMA).
the
with
ossified
voor
Museum,
morphometric
largely
are
specimens
rioventrally
all
taking
Isbriicker
by
for
of
Instituut
Zoologisch
Amsterdam
van
data
proposed 182);
Zoologie,
methods
meristic
the
(MBUCV) and of
Taxonomische
have
I,
five 4, i
have
rays,
I, 5 the
to its base.
—
Dentectus is from the Latin dens
tooth,
and
covered,
allusion to the
from
the
concealed,
Latin
tectus
disguised,
an
unique dentition.
127
Comparison Dentectus
related
the
on
with
hand
one
seems
Pseudohemiodon &
Rhadinoloricaria Isbriicker Crossoloricaria
Isbriicker,
to
be
closely
Bleeker,
Nijssen,
various
morphometric
In
the
of
measured
1862,
latter
62.5
show
and
Dentectus
and
141.6
the
in
characters.
standard
specimens
of
mm,
differences
meristic
following comparison,
1974, and
(these
1979
Planiloricaria
and
ra,
are
various
lengths
between of
species
~
the three
within
genera
defined
Pseudohemiodonof group
the
constitute
Pseudohemiodongroup genera
the
subtribe
by Isbriicker,
Loricariina,
On the other
1981).
as
hand,
lengths
are
known
between
specimen
Dentectus
it
also
possesses
below) pointing caria of
to
Isbriicker,
the
subtribe
characters
some
a
(indicated
relationship with Planilori-
1971,
hitherto the
only
3.1-4.0 times in and
6.6,
SL,
1979)shares
genera: head
Pseudohemiodon-group depressed, more or
caria
broad
the snout
teeth
acute),
in
dorsal
less disc-like to
has
presence
of
notch),
dorsally large
rays
and often with
Dentectus,
128
1.
size
of or
head
and
in
(some
small
a
the
profile
of
number,
species a
of
pupil,
snout,
a
rather
ridges outer
caudal fin
in
on
extremely long filament.
barbarmatus
n.
gen.
et
n.
sp.,
ray in
al.,
0.4-6.9
genera, and caudal
Dentectus
the
contained
4.2-5.1
in
al.,
Pseudohemiodon et
and
fin
against
5.5
Dentectus
in SL
'spine' is 0.3-0.5
in
0.6
Pseudohemiodon in
6.1-9.9
and
Planiloricaria. The
is
'spine'
1.6-4.7
in
'n
SL
in
Pseudohemiodon et
in Planiloricaria.
has the maximum orbital diameter
contained 5.6-6.4 times
Pseudohemiodon-group
in head
it is
length,
4.3-7-2,
in
the
in Planilori-
caria 13.1. The interorbital width of Dentectus is
(un-
produced
5.1-7.3
Dentectus,
lower
in
head
al.,
the Pseudohemiodonof genegroup
Dentectus
4.5first
has
in Planiloricaria.
5.5
related
posterior
flap extending
prominent
and
'spines'
an
snout
contained
It
2.4.
The upper caudal fin
very much
anterior and
rather
fold,
dorsal surface
Fig.
usually
the
tapering, relatively
notch
downward from
labial
branched)
an
with
triangular (Rhadinolori-
in
orbital
Pseudohemiodon have orbital
view,
profile
reduced
an
characters
most
214
spine
Pseudohemiodon et al.
fin
dorsal
against
and Dentectus
SL,
fin
only
mm.
genus
Isbriicker,
(cf.
the dorsal
standard
mm; the
of Planiloricaria is
Planiloricaria
branched
Planiloricariina
has
63 and 246.5
the
holotype in
and
length, 5.2
in
4.2-4.6
Pseudohemiodon et
in Planiloricaria.
Dentectus
lateral
4.3-6.1
view.
has
16-18
coalescing
lateral
body
Fig.
2.
dorsal
Dentectus
barbarmatus
n.
gen.
et
n.
sp.,
details
of anterior
part
of
holotype
in
ventral
(left half)
and
in
(right half) view, respectively.
129
Pseudohemiodon
scutes,
Planiloricaria
There
19-20.
scutes
body
in
al.
et
are
Dentectus,
hemiodon-group
13-21,
lateral
31-33 in
31-34
and
genera,
have
in
40
Pseudo-
Isbrlicker,
pointed
specimens
teeth.
visible
teeth
in
dentaries
simply
the to
proved
be
in-
have
staining,
and
the
in
Neither
present.
speci-
Pseudohemiodongroup genera,
of Planiloricaria have been cleared and
the
the
and
the former
counted in the premaxilla, The
dentary.
to
genus-group, up
lack
of
a
numerous
only
and up
to
visible
P.
like
dentary,
The
in
11
(figs.
the
hand
at
has
Dentectus.
Holotype,
presence
scutelets) lips,
and
jaws
absent
characters,
there
However, various
teeth
Dentectus
all
in
are
the
to
three
As
appropriate
subtribe
to be the best
of
and
in
and
by
along the
all
the
and
buccal
previously),
barbels
Linnaeus,
130
by
now
the
seems
Pseudo-
Planiloricaria For
the
expanded
this
of Rio
the
of
(small
of
and
cavity.
and
of
(two
long
lower
(in
also
a
more
relatively
a
few
the
from the
large
teeth;
genera
Brochiloricaria
L.
ZMA
116.648,
Estado Sum
J.
(o8°4i' N,
N.
W), 0.
Baskin,
paratype,
Silva
Puente Proven-
31-VII-1981; SL
mm,
coll.
& L.
Estado
from
upriver coll.
F.
89.1
mm,
Venezuela,
F.
Mago Leccia,
F.
2-VIII-
Aguana,
MBUCV-v-12940,
Venezuela,
mm,
mm,
Venezuela, the
coll.
one,
88.9
99
MBUCV-
Rio Bocono at Puerto Sum
Portuguesa,
69°i2' W), L.
SL
one,
at
Aguana,
paratypes, MBUCV-v-13173,
SL
one,
Portuguesa,
Guanarito
Rio
(o8°42' N,
0. Castillo
Provenzano,
&
Aguana, 28-VII-1981.
Description.— Morphometric
possessing
&
and
mm,
Masparro
116.638,
holotype
as
paratypes,
N, 69°38' W),
(o8°2i'
Fernandez MBUCV-v-
ZMA
59.5
a
Baul
lips
sexual
restricted
SL
El
of
data
140.8-141.6 Rio
0. Castillo
zano,
N.E.
mm,
mm,
Salinas,
Agustin
same
paratypes,
SL
136.5
Rio
paratypes,
mm,
Barinas,
Libertad
coll.
—
stained);—
two,
v-12920, Estado
SL
Viejo,
59.5-120.5
136.8
and
Guanare,
few
SL
these
cleared
67.3
and
Secondary
easily distinguished
(comprising 1758,
SL
one,
Pao
25-II-1950;
small
numerous
upper
—
Cojedes,
68°oy' W),
12985, six,
char-
are
possession
presence
on
subtribe Loricariina
Loricariina
mutual
is unknown.
The subtribe is
of
These
Yepez,
1974;
reduced dentition
a
dimorphism
than
and
as
Pseudo-
place.
acterized above
teeth),
it
Loricariini.
The Planiloricariina thus
barbels
genus
the
close
Dentectus,
genera,
and
the subtribe Planiloricariina appears
assignment
with
in this
consequence,
of
hemiodon-group
Loricariidae.
similarities
rather
the
unique
Planiloricaria.
a
assign
to
single
a
(small
are
genera of
to
indicate
relationship.
jaws
other
many
and
hemiodon-group similarities
a
ossifications
overlapping characters
compared
to
of
table 1)
(fig. 6).
Estado
Venezuela,
and the structure of
barbels,
on
1-5;
MBUCV-v-12780,
(09°i3' N,
dermal
species
new
no
position of
has
1979
1981).
tributary Systematic
Isbriicker,
(cf.
were
whereas it has 3 teeth
premaxillary teeth,
in each
by
subtribe
of Planiloricaria
specimen
cryptodon)
The
nor
Material examined
of
(holotype
differ
others
branched barbels,
dentition
well-developed
and
Two
premaxillae
among
barbels.
conspicuous
Ricolina with
usually
jaws.
stained.
teeth
9
the
Hemiodontichthyina)
Planiloricariina
Dentectus barbarmatus In
of
preserved
and
premaxillae
firm and
are
one
from
barbarmatus
Only after clearing
both
of the
mens
above,
out
Dentectus
of
in
or
subtribes with invisible teeth in the
(Reganellina
As
and Paraloricaria
1979,
the teeth
1979)
in both
long
very
Isbriicker,
Planilori-
caria.
3
in
Nijssen,
sense
in
some
number in
the
Loricaria
Isbriicker
&
table I. of
the
Dermal moderate on
the
scutes.
and meristic
Illustrations of
paratypes
ossifications, odontodes,
coalescing Two
are
on
given
in
and
are more
the parallel the
presented
are
holotype
fin-spines
which
and
ridges
data the
and
figs. rays
of
1-5. with
prominent
lateral
supraoccipital
body and
Fig.
on
Dentectus
3.
the
ridge
two
on
nuchal
adjacent
the
plate
larger
by
third
with
supraoccipital. a
barbarmatus
a
a
gen.
predorsal predorsal ridge
Adjacent
scute with
n.
at
to
a
n.
sp.,
scutes,
a
A
scute.
either side
the nuchal
ridge, posteriorly
6 anterio-dorsolateral scutes,
with
et
paratype
single small of the
plate
is
continued
each of
which
ridge.
Orbital
rim
in
ZMA
small and more
oval,
a
conspicuous
though
in
dorsal,
shallow
prominent
Dorsal
surface
lateral
posterior
diverges
of
in front of
parallel ridges
the
At
and
notch
ventral
and
head
with
a
a
to
near
is
another
the
ridge,
along
This
the nostrils into two
extending
rim
with
ridge
areas.
either side between these
orbital
view.
anterior notch.
the interorbital and internasal
snout.
with
116.638
tip
ridge
closely of the
ridges being
and an
131
ventral in
1 6. 38 ZMA in
par type of
part
anterior of
details sp.,
etn. gen. n.
view.
bar matus (right) Dent cus
dorsal in
and 4.
Fig. (left)
132
Fig.
Dentectus
5.
details
of
position
of
the
ridges;
anterior one
one
it
anteriorly
margin
of the
diverges stretching
anterior border of converge and
Ventral surface
marginal series lets.
—
(d)
distal
part
supraoccipital
orbital
into
posterior
rim.
two
beyond
the other
At
the
branches, the
outer
embracing
the
continuing
to
with the first branch.
of head
naked,
of somewhat
half
— (a) right
MBUCV-v-12985,
of the
the orbit and
merge
in
embracing the nostril,
the
nostrils,
the
sp.;
et n.
gen.
paratype
height
branch
bordering
edge
of
at the
diverges
other
n.
stained
jaws; — (c) premaxilla;
orbital notch, the
and
continuation
anterior it
barbarmatus
cleared
a
except
rectangular
of
SL
for
a
head
99
mm;
of
—
in
holotype (b)
central
ventral area
of
view; — (b-d)
mouth, showing
dentary.
Preanal
the
series,
scutelets. from
the
merges
ing
the
covered
area
scutelets,
shaped
This
a
by
height
a
small
into
irregularly
of
consisting
of scutelets is naked
narrow
much wider naked
irregular
7
smaller
separated
strip,
area
which
surround-
pelvic fins.
irregular
decrease
riorly
series
complex
anus
into
with
arranged
outer
Abdominal and rather
scute-
of
in
size
of the there
thoracic
pattern
areas
of
anteriorly,
branchiostegal
are
5
covered
scutelets,
reaching
by
a
which to
the
membrane. Poste-
to 9 scutelets in
a
transverse
133
Table of
1. — Morphometric
mm;
holotype in
in
—
MBUCV-v-12985; data of
H,
characters indicate
4-17
are
scattered
widely
—
paratype
identical
A,
in
and
F,
ratios
of
as
a
SL,
in
the
standard
axial length
Dentectus data of
B,
barbarmatus
paratype
paratype in MBUCV-v-12920;
characters
data of
18-35
paratype ratios
are
adjacent column;
in
of
asterisk in
n.
gen.
in ZMA
—
G,
et
n.
116.638;
data of
sp.;
—
paratype
HL,
and
in
(A),
characters
H indicates
of
3
paratypes
MBUCV-v-13173;
41-44 the
measurements
data
Characters
MBUCV-v-12940.
column
—
C-E,
are
high
1-3
are
counts.
number
in
—
mm,
Arrows
of
small,
scutelets.
(A)
2
of
—
116.648; — J,
specimen
I
data
holotype;
data of
ZMA
figures
meristic
data of
length
136.5 —
A
136.5
B
C
136.8
120.5
150.6
—
—
D
E
F
92-3
62.5
141.6
—
—
—
G
89.1 —
II
88.9 99-9
.1
67-3 —
total
length
493-5
493-5
200.5
362.3
4
head
length
29.3
4-7
4.8
4-8
4-4
4-2
4.6
4-3
4-i
4.2
5
predorsal length
43-5
3-1
3-2
3-2
3.0
2.9
3-i
2.9
-■')
30
6
postdorsal length
78.1
i-7
i-7
1-7
i-7
1.8
1.7
1.8
1.8
1.8
7
preanal
65.0
2.1
2.2
2.2
2.1
2.1
2.1
2.0
2.0
2.1
2.0
3
length
8
postanal
9
distance snout to
length
io
distance snout to
11
dorsal
12
first
dorsal
anal
spine length
13
anus
pelvic
spine length length
ray
14
pectoral spine length
15
pelvic spine length
16
upper
caudal
17
lower
caudal
18
snout
length
lower
lip
19 20
21
22
23
24
lower
lip
maxillary thoracic
spine length spine length
66.5
2.1
50.4
2-7
42.6
abdominal maximum
178.5
507-1
2.1
2.1
2.1
2.8
2.7
2.6
2.6
2-7
—
267.3
2.2
2.2
2.5
24 2.8
3-i
3-0
3-1
3-1
